Eastern Mojave Vegetation | Images -- Species - Poaceae |
By Tom Schweich |
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Images are organized by the topics listed at left. Click on a topic name to see images with that topic. Would you like to use these photographs in your school term paper or report? License is hereby granted for non-commercial use of the images contained herein provided that the images are reproduced with the copyright notice intact. This means that we give you our permission to use the photographs and information you find here in your school term papers and reports. You may resize the photograph to fit your needs, but you may not remove the copyright. Your term paper should note that the photographs are reproduced with permission and it should have a reference to this web site, such as: "Photograph ©1999 Tom Schweich. Used by permission, http://www.schweich.com" If you have a comment or a question about a photograph you may write to me at tomas@schweich.com I will answer you as soon as I can. I sometimes post interesting questions in my FAQ, but I will not disclose your full name or address.
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Habitat of three common grasses in Windy Saddle Park | Photographed 24 July 2015. The three grasses are: Bouteloua gracilis, B. curtipendula, and Aristida purpurea.
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| Aegilops cylindrica
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Inflorescence of Coll. No. 1139, Aegilops cylindrica | Scanned 14 October 2015.
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Inflorescence of Coll. No. 1139, Aegilops cylindrica | Scanned 14 October 2015.
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| Agropyron cristatum
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Coll. No. 3080, | Photographed 31 December 2023.
Coll. No. 3080, 12 July 2023, characters observed: Perennial grass, to 75 cm., not obviously rhizomatous; Leaves, sheath, 93 mm., open, ligule, membranous, auricles, minimal, 0.5 mm., lanceolate, blade, 160 mm. × 3.2 mm. wide, flat, adaxial pilose; Inflorescence, 2-sided spike, 45 mm. × 7 mm. wide; Spikelets, many, 12 mm. × 12 mm. wide, overlapping about ⅔ or 8 mm., similar, borne singly, glumes and florets spreading; Glumes, lower, 5 mm. + awn, 3 mm. Veins, 5, upper, 5 mm. × 1.5 mm. wide + awn 3.5 mm., <lowest floret, veins, 5, keeled; Florets, 5 per spikelet; Lemma, 7.5 mm., >glumes, keeled, barely scabrous on veins, veins, 5, tip, bifid, awns, #1, 2.8 mm., straight; Palea, 5.8 mm., <lemma, membranous, keel, ciliate, tip, bifid; Anthers, 3 mm.
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My collection of Agropyron cristatum at Denver Botanic Gardens | ©reserved Denver Botanic Gardens URL: http://swbiodiversity.org/seinet/collections/individual/index.php?occid=21435268&clid=0 Accessed 12 April 2020. Usage Rights: CC0 1.0 (Public-domain)
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Spikelet of Coll. No. 3080, | Photographed 31 December 2023.
Coll. No. 3080, 12 July 2023, characters observed: Perennial grass, to 75 cm., not obviously rhizomatous; Leaves, sheath, 93 mm., open, ligule, membranous, auricles, minimal, 0.5 mm., lanceolate, blade, 160 mm. × 3.2 mm. wide, flat, adaxial pilose; Inflorescence, 2-sided spike, 45 mm. × 7 mm. wide; Spikelets, many, 12 mm. × 12 mm. wide, overlapping about ⅔ or 8 mm., similar, borne singly, glumes and florets spreading; Glumes, lower, 5 mm. + awn, 3 mm. Veins, 5, upper, 5 mm. × 1.5 mm. wide + awn 3.5 mm., <lowest floret, veins, 5, keeled; Florets, 5 per spikelet; Lemma, 7.5 mm., >glumes, keeled, barely scabrous on veins, veins, 5, tip, bifid, awns, #1, 2.8 mm., straight; Palea, 5.8 mm., <lemma, membranous, keel, ciliate, tip, bifid; Anthers, 3 mm.
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Coll. No. 821, Agropyron cristatum, "Crested Wheat Grass" | Photographed 30 May 2012 … although, the infloresence is much tighter than the illustration in TJM2 would suggest.
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Habitat of Agropyron cristatum near June Lake. | Photographed 30 May 2012.
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Coll. No. 821, Agropyron cristatum | Photographed 26 January 2013.
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| Agrostis gigantea
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Coll. No. 3211, | Photographed 1 February 2024.
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My collection of Agrostis gigantea from Ranson/Edwards. | ©reserved Denver Botanic Gardens URL: http://swbiodiversity.org/seinet/collections/individual/index.php?occid=8550168&clid=0 Accessed 12 April 2020. Usage Rights: CC0 1.0 (Public-domain).
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E. H. Brunquist collection of Agrostis gigantea | ©reserved Denver Botanic Gardens URL: http://swbiodiversity.org/seinet/collections/individual/index.php?occid=3058218&clid=0 Accessed 12 April 2020. Usage Rights: CC0 1.0 (Public-domain). I wanted to use this image because it shows a strong rhizome.
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| Agrostis scabra
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Coll. No. 2700.1, Agrostis scabra. | Photographed 10 February 2022.
Coll. No. 2700.1, 17 Aug 2021, characters observed: Perennial grass, 40 cm., rhizomes not seen; Leaves, sheath, 30 mm., open, distally retrorsely scabrous, ligule, membranous, 2 mm., auricles, absent, blade, 35 mm. × 0.8 mm. wide, flat, margins, scabrous; Inflorescence, 1 per culm, >leaves, open panicle with ascending branches, 140 mm. × 80 mm. wide; Rachis, not extended; Spikelets, many, 2 mm. × 0.6 mm. wide, similar, borne singly, pedicels 1.7-3.5 mm, scabrous; Compression, lateral; Disarticulation, above the glumes; Glumes, lower, 2.5 mm., keeled, scabrous on keel, veins, #1, upper, 2.4 mm., >lowest floret, keeled, scabrous on keel, veins, 1, awns, 0; Florets, 1 per spikelet, fertile; Lemma, 1.5 mm., <glumes, rolled, glabrous, translucent, veins, ?, tip, obtuse, awns, 0; Palea, 0.3 mm., membranous, colorless; Anthers, 0.5 mm.; Achene, 1 mm., green, glabrous.
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Coll. No. 3124, Agrostis scabra. | Photographed 14 January 2024.
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| Agrostis stolonifera
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Coll. No. 514, Agrostis stolonifera | Scanned 17 March 2011. Background grid is 10 x 10 to the inch.
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Coll. No. 1225, Agrostis stolonifera | Scanned 14 December 2015. Coll. No. 1225, 16 Jul 2015, characters observed: Perennial grass, to 1 m., no sign of either rhizomes or stolons; Stem, nodes green; Leaves, sheath, open, glabrous at summit, ligule, membraneous, 4 mm., angled, auricles, 0 (double fold of proximal blade margin), blade, 85 mm. × 3.8 mm. wide, scabrous abaxially; Inflorescence, open panicle, 18 cm. × 14 cm. wide, deltate; Spikelets, 1 per node, on 1°-3° panicle branches, axis not extended beside or above floret; Glumes, #2, 2.2 mm., ±equal, keeled, scabrous on keel, veins 1, awns 0, purplish; Disarticulation above the glumes; Florets, 1 per spikelet, bisexual; Lemma, 1.8 mm., transparent, except 3 green veins, awns 0; Palea, 1.5 mm.; Anthers, #3, 1.4 mm., tan with purple abaxially.
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Roots of Coll. No. 1515, Agrostis stolonifera | Scanned 12 January 2017. Ranson/Edwards Homestead Open Space Park, Jefferson County, Colorado. Northern part of open space, 1.6 mi. west on CO Highway 72 from CO Highway 93, north 1.9 mi. on Plainview Road, then 300 m. northeast, almost to the boundary fence; 16.3 km. north northwest of the GNIS location of Golden. 39.8991°N, 105.2668°W. WGS 1984 Elev. 1937 m. Weakly stoloniferous, growing in and beside ephemeral stream. Introduced. Coll. No. 1515, 24 July 2016, characters observed: Perennial grass, to 85 cm., decumbent at base, weakly rhizomatous, 1 of 3 collections has a 2.5 cm. scaly rhizome; Leaves, sheath, open, ligule 4 mm., chartaceous tissue extending down sheath 20 mm., auricles, 1 mm., blade, 80 mm. × 3.2 mm. wide, flat, scabrous adaxially; Inflorescence, panicle, branches not bearing spikelets to base; Spikelets, 1 floret; Glumes, ±equal, 2.3 mm., purplish, keeled scabrous; Lemma, 1.8 mm. + awn 0.3 mm., thin, translucent, veins, 3(5?); Palea, 1.3 mm. Introduced.
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Coll. No. 1515, Agrostis stolonifera | Photographed 12 January 2017. Ranson/Edwards Homestead Open Space Park, Jefferson County, Colorado. Northern part of open space, 1.6 mi. west on CO Highway 72 from CO Highway 93, north 1.9 mi. on Plainview Road, then 300 m. northeast, almost to the boundary fence; 16.3 km. north northwest of the GNIS location of Golden. 39.8991°N, 105.2668°W. WGS 1984 Elev. 1937 m. Weakly stoloniferous, growing in and beside ephemeral stream. Introduced. Coll. No. 1515, 24 July 2016, characters observed: Perennial grass, to 85 cm., decumbent at base, weakly rhizomatous, 1 of 3 collections has a 2.5 cm. scaly rhizome; Leaves, sheath, open, ligule 4 mm., chartaceous tissue extending down sheath 20 mm., auricles, 1 mm., blade, 80 mm. × 3.2 mm. wide, flat, scabrous adaxially; Inflorescence, panicle, branches not bearing spikelets to base; Spikelets, 1 floret; Glumes, ±equal, 2.3 mm., purplish, keeled scabrous; Lemma, 1.8 mm. + awn 0.3 mm., thin, translucent, veins, 3(5?); Palea, 1.3 mm. Introduced.
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| Alopecurus arundinaceus
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Schweich collection of Alopecurus arundinaceus | ©reserved Denver Botanic Gardens URL: http://swbiodiversity.org/seinet/collections/individual/index.php?occid=13288074&clid=0 Accessed 23 April 2020. Usage Rights: CC0 1.0 (Public-domain).
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Coll. No. 2274.1, Alopecurus arundinaceus. | Photographed 28 October 2020.
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| Alopecurus carolinianus
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Coll. No. 1689.3, Alopecurus carolinianus | Scanned 28 June 2017. Orginally determined A. geniculatus, but the small size of the anthers is more suggestive of A. carolinianus.
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Inflorescence of Coll. No. 1689.3, Alopecurus carolinianus | Scanned 25 December 2017. I originally determined this A. geniculatus. The anthers are quite small, less than 1 mm., as can be seen on the lower left side of the inflorescence. The small length of the anthers was first noted by Janet Wingate. This character is more suggestive of A. carolinianus.
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Coll. No. 1689.3, Alopecurus carolinianus | Scanned 25 December 2017. Orginally determined A. geniculatus, but the small size of the anthers is more suggestive of A. carolinianus.
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| Alopecurus geniculatus
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Coll. No. 1930.2, Alopecurus geniculatus | Photographed 11 December 2018. Coll. No. 1930.2, 8 Jun 2018, characters observed: Perennial grass, clumped, 45-50 cm.; Roots, fibrous; Leaves, sheath and blade differentiated; Sheath, 40-55 mm., open, loose or inflated; Ligule, membranous, 2.5-7.0 mm., auricles, 0; Blade, 65-75 mm. × 2.2 mm. wide, flat, glabrous; Inflorescence, 1 per culm, >leaves, panicle tightly contracted, 50 mm. × 6 mm. wide, branched ≥ 2×, spikelets, many; Rachis, not extended, glabrous; Spikelets, many, similar, borne singly, 1-flowered, 3 mm. × 1.3 mm. wide, look like little horseshoe crabs; Compression, lateral; Disarticulation, below the glumes; Glumes, lower, 2.8 mm., upper, 2.7 mm., keeled, membranous, awns, 0, hairy on keel and lateral vein; Florets, 1 per spikelet, bisexual; Lemma, 2.7 mm., membranous, glabrous, awns, #1, 5 mm., gently curved not sharply bent, attached proximal ⅓; Palea, absent; By dimensions of glumes, lemmas, and lemma awns, this would be A. geniculatus, but very few awns geniculate.
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Schweich collection of Alopecurus geniculatus | ©reserved Denver Botanic Gardens URL: http://swbiodiversity.org/seinet/collections/individual/index.php?occid=21435339&clid=0 Accessed 24 April 2020. Usage Rights: CC0 1.0 (Public-domain).
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| Andropogon gerardii
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Dissected spikelet pair of Coll. No. 1060, Andropogon gerardii | Photographed 4 June 2015. Coll. No. 1060, 28 Aug 2014, characters observed: Perennial, to 85 cm., clumped, rhizomes unknown; Leaves, blade and sheath differentiated; Sheath, 11 cm., open; Ligule, membraneous (though surrounded by hairs), 3 mm., auricles 0; Blade, soft, flat, 27 cm., hairy near ligule; Inflorescence, 80 mm., > leaves, 1-2 per stem, in 5-6 digitate branches, spikelets many; Rachis, soft bristles on margins and at nodes; Disarticulation at nodes of rachis; Spikelets in pairs, Sterile spikelet, 7 mm., pedicel 4.5 mm.; Fertile spikelet, Florets #1, Glumes, awns 0, ±equal, lower keeled, upper flattened and grooved; Lemma, 6 mm., membraneous, rolled, glabrous, tip 2-forked, awn, #1, 13 mm., bent, twisted, attached on back; Stamens, 3; Anthers, yellow; Ovary, glabrous; Stigma, feathery.
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Spikelets of Coll. No. 1265, Andropogon gerardii | Scanned 7 January 2015.
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Inflorescence of Coll. No. 1265, Andropogon gerardii | Scanned 7 January 2015.
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| Andropogon hallii
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Coll. No. 1060, Andropogon hallii | Photographed 28 August 2014.
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| Apera interrupta
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Coll. No. 2547, Apera interrupta | Photographed 28 November 2021.
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| Aristida purpurea
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Coll. No. 1140, Aristida purpurea | Photographed 9 June 2015.
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| Aristida purpurea longiseta
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Coll. No. 1140, Aristida purpurea var. longiseta | Scanned 15 October 2015. Coll. No. 1140, 9 Jun 2015, characters observed: Perennial grass, 20 cm. when collected, possibly elongating in maturity; Inflorescence, panicle, branches not pulvinate, i.e., no swelling at bases of pedicels; Axis ±straight; Pedicels, stiff, ±straight; Spikelets with 1 floret; Glumes, 2, lower, 11 mm., veins 1, upper, 20 mm.; Lemma, 11 mm., smoothly transitioning to awns without twisted column, apex 0.4-0.6 mm. wide, awns, 3, 58 mm., straight. Determined var. longiseta per length of upper glumes and dimensions of lemma.
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| Arrhenatherum elatius
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Coll. No. 2442, Arrhenatherum elatius | Photographed 31 December 2020,
Coll. No. 2442, 28 August 2020, characters observed: Perennial, to 115 cm., tufted, no rhizomes observed; Leaves, sheath, 100 mm., open, ligule, 2.5 mm., membranous, erose, blade, 30 mm. × 5 mm. wide, flat, barely scabrous; Inflorescence, 1 per culm, >leaves, panicle, 200 mm. × 40 mm. wide; Rachis, not extended, delicate; Spikelets, many, 8 mm. × 2.3 mm. wide + awn, similar, borne singly; Disarticulation, above the glumes; Glumes, very thin, translucent, lower, 5 mm., veins, #1, tip bifid, upper, 9 mm., =lowest floret, veins, #3, ±parallel, awns, absent; Florets, 2 per spikelet; Callus, straight hairs, 2.3 mm.; Lemma, 8 mm., =glumes, veins, #7, awn, #1, 14 mm., twisted, round, attached near base (2.3/8.0); Palea, 6 mm., membranous, margin, ciliate.
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| Arundo donax
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Max Lichter photograph of Arundo donax | ©Max Lichter (CC BY-SA). Locality: near Red Rock Crossing, Sedona, Arizona URL: https://serv.biokic.asu.edu/imglib/h_seinet/seinet/Poaceae/Arundo-donax-P-web-9-23-1.jpg Accessed: 10 April 2020. Images are made available on SEINet under the Creative Commons Attribution-ShareAlike (CC BY-SA).
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| Avena fatua
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Coll. No. 812, Avena fatua | Scanned 15 Jun 2013. Coll. No. 812, 8 Apr 2012, characters observed while keying: Annual, to 100 cm; Roots fibrous; Stem herbaceous; Internodes hollow; Leaf blade and sheath differentiated; Sheath open; Blade soft, margins glabrous; Inflorescence many, panicle, > leaves; Spikelets many on 1° and 2° branches, 20-25 mm.; Florets 3 per spikelet; Glumes present, ±equal, veins 9, awns 0; Axis hairy on abaxial side; Callus bearded; Lemma 10-12 mm, rounded, veins >= 5, tip 2-forked, awns 1, attached at middle, 15 + 25 = 40 mm, single bent, twisted below, straightening when wetted; Palea, ±< lemma, glabrous, margins ciliate; Ovary hairy.
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| Blepharoneuron tricholepis
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Blepharoneuron tricholepis (Torr.) Nash | Scanned 26 September 2014.
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Blepharoneuron tricholepis (Torr.) Nash | Scanned 26 September 2014.
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Blepharoneuron tricholepis (Torr.) Nash | Scanned 26 September 2014.
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| Bouteloua curtipendula
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Coll. No. 1241, Bouteloua curtipendula | Photographed 24 July 2015.
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Coll. No. 1241, Bouteloua curtipendula | Scanned 20 December 2015.
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| Bouteloua gracilis
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Coll. No. 3146, Bouteloua gracilis. | Photographed 17 January 2024.
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Habitat of Coll. No. 1227, Bouteloua gracilis | Photographed 20 July 2015.
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Coll. No. 1242, Bouteloua gracilis | Scanned 20 December 2015.
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| Bromus briziformis
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Habit of Coll. No. 2167, Bromus briziformis | Photographed 21 July 2019.
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Dissected spikelet of Coll. No. 2167, Bromus briziformis | Scanned 22 July 2019. Left to right: lower glume, floret showing lemma, achene, and palea, then upper glume, and outside of floret. The main this to see is the number of veins, which I count as nine on the upper glume and the lemma, and that there are not uniformly spaced. Coll. No. 2167, 21 July 2019, characters observed: Annual grass, 30-40 cm., clumped; Roots, fibrous; Stem, herbaceous; Leaves, blade and sheath differentiated, soft; Sheath, closed; Ligule, membraneous, ciliate; Blade, 120 mm. × 4 mm. wide, flat, hairy; Inflorescence, 1 per culm, nodding, raceme or few-branched panicle, >leaves; Rachis, not extended, scabrous; Spikelets, many, 16-20 mm. × 8-10 mm. wide, similar, borne singly, pedicels, 18-22 mm.; Compression, lateral; Disarticulation, unknown, rachilla seems pretty tough; Glumes, spreading at 45° to rachilla, lower, 5.5 mm., veins, #5, terete, upper, 7.0 mm., <lowest floret, veins, #9, ±parallel, membraneous, tough, awns, #0; Florets, 10-12 per spikelet, spreading at 45° to rachilla; Lemma, 8 mm., >glumes, ovate, membraneous, rolled, glabrous, tan, veins, #9, not uniformly spaced, ±parallel, margins, scarious, tip acute, awns, #0; Palea, 6 mm., membraneous, ciliate, tip, unknown, color, tan; Stamens, none seen; Achene, 5 mm., brown, fused to lemma and palea.
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Spikelet of Coll. No. 2167, Bromus briziformis | Scanned 22 July 2019. Coll. No. 2167, 21 July 2019, characters observed: Annual grass, 30-40 cm., clumped; Roots, fibrous; Stem, herbaceous; Leaves, blade and sheath differentiated, soft; Sheath, closed; Ligule, membraneous, ciliate; Blade, 120 mm. × 4 mm. wide, flat, hairy; Inflorescence, 1 per culm, nodding, raceme or few-branched panicle, >leaves; Rachis, not extended, scabrous; Spikelets, many, 16-20 mm. × 8-10 mm. wide, similar, borne singly, pedicels, 18-22 mm.; Compression, lateral; Disarticulation, unknown, rachilla seems pretty tough; Glumes, spreading at 45° to rachilla, lower, 5.5 mm., veins, #5, terete, upper, 7.0 mm., <lowest floret, veins, #9, ±parallel, membraneous, tough, awns, #0; Florets, 10-12 per spikelet, spreading at 45° to rachilla; Lemma, 8 mm., >glumes, ovate, membraneous, rolled, glabrous, tan, veins, #9, not uniformly spaced, ±parallel, margins, scarious, tip acute, awns, #0; Palea, 6 mm., membraneous, ciliate, tip, unknown, color, tan; Stamens, none seen; Achene, 5 mm., brown, fused to lemma and palea.
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| Bromus carinatus
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Spikelets of Coll. No. 2090, Bromus carinatus | Scanned 21 October 2019. Coll. No. 2090, 26 June 2019, characters observed: Annual/Perennial unknown (planted Spring 2019), height 60-90 cm., clumped; Leaves, sheath and blade differentiated, soft; Sheath, 80 mm., closed, generally glabrous, except collar somewhat hairy; Ligule, membraneous, 2 mm.; Auricles, absent; Blade, 140 mm. × 7 mm. wide, flat, scabrous, mid-rib prominent; Inflorescence, 1 per culm, >leaves, panicle, 180-200 mm. × 40-90 mm. wide; Rachis, not extended, minutely scabrous on abaxial side; Spikelets, many (#22-26), 36 mm. × 11 mm. wide, similar, 1 per node; Compression, lateral; Disarticulation, between the florets; Glumes, lower, 9.5 mm., veins, #5, distinct, upper, 12 mm., <lowest floret, veins, #7, distinct keeled, membraneous, awns, 0, margins short ciliate; Florets, 6 per spikelet; Rachilla, exposed, scabrous on abaxial side, not extended; Lemma, 13 mm., broad, 2.5 mm. midvein to margin, >glumes,ovate, keeled distally, finely short hairy throughout, veins, #9, glabrous, margins, scarious, tip, entire, awns, #1, 8 mm., straight, attached near tip; Palea, 7 mm., <lemma, keel, ciliate, veins, green, margin, glabrous, tip, entire; Anthers, 3.6 mm., yellow; Ovary, hairy. Keys poorly in most Colorado floras, but quite nicely to B. carinatus var. carinatus in the Jepson Manual, in good part by the length of the lemma awn. Regardless, I think this is very likely a commercial hybrid used in revegetation.
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Inflorescence of Coll. No. 2090, Bromus carinatus | Scanned 21 October 2019. Coll. No. 2090, 26 June 2019, characters observed: Annual/Perennial unknown (planted Spring 2019), height 60-90 cm., clumped; Leaves, sheath and blade differentiated, soft; Sheath, 80 mm., closed, generally glabrous, except collar somewhat hairy; Ligule, membraneous, 2 mm.; Auricles, absent; Blade, 140 mm. × 7 mm. wide, flat, scabrous, mid-rib prominent; Inflorescence, 1 per culm, >leaves, panicle, 180-200 mm. × 40-90 mm. wide; Rachis, not extended, minutely scabrous on abaxial side; Spikelets, many (#22-26), 36 mm. × 11 mm. wide, similar, 1 per node; Compression, lateral; Disarticulation, between the florets; Glumes, lower, 9.5 mm., veins, #5, distinct, upper, 12 mm., <lowest floret, veins, #7, distinct keeled, membraneous, awns, 0, margins short ciliate; Florets, 6 per spikelet; Rachilla, exposed, scabrous on abaxial side, not extended; Lemma, 13 mm., broad, 2.5 mm. midvein to margin, >glumes,ovate, keeled distally, finely short hairy throughout, veins, #9, glabrous, margins, scarious, tip, entire, awns, #1, 8 mm., straight, attached near tip; Palea, 7 mm., <lemma, keel, ciliate, veins, green, margin, glabrous, tip, entire; Anthers, 3.6 mm., yellow; Ovary, hairy. Keys poorly in most Colorado floras, but quite nicely to B. carinatus var. carinatus in the Jepson Manual, in good part by the length of the lemma awn. Regardless, I think this is very likely a commercial hybrid used in revegetation.
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Coll. No. 2090, Bromus carinatus | Photographed 21 October 2019. Coll. No. 2090, 26 June 2019, characters observed: Annual/Perennial unknown (planted Spring 2019), height 60-90 cm., clumped; Leaves, sheath and blade differentiated, soft; Sheath, 80 mm., closed, generally glabrous, except collar somewhat hairy; Ligule, membraneous, 2 mm.; Auricles, absent; Blade, 140 mm. × 7 mm. wide, flat, scabrous, mid-rib prominent; Inflorescence, 1 per culm, >leaves, panicle, 180-200 mm. × 40-90 mm. wide; Rachis, not extended, minutely scabrous on abaxial side; Spikelets, many (#22-26), 36 mm. × 11 mm. wide, similar, 1 per node; Compression, lateral; Disarticulation, between the florets; Glumes, lower, 9.5 mm., veins, #5, distinct, upper, 12 mm., <lowest floret, veins, #7, distinct keeled, membraneous, awns, 0, margins short ciliate; Florets, 6 per spikelet; Rachilla, exposed, scabrous on abaxial side, not extended; Lemma, 13 mm., broad, 2.5 mm. midvein to margin, >glumes,ovate, keeled distally, finely short hairy throughout, veins, #9, glabrous, margins, scarious, tip, entire, awns, #1, 8 mm., straight, attached near tip; Palea, 7 mm., <lemma, keel, ciliate, veins, green, margin, glabrous, tip, entire; Anthers, 3.6 mm., yellow; Ovary, hairy. Keys poorly in most Colorado floras, but quite nicely to B. carinatus var. carinatus in the Jepson Manual, in good part by the length of the lemma awn. Regardless, I think this is very likely a commercial hybrid used in revegetation.
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| Bromus ciliatus
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Coll. No. 1993, Bromus ciliatus | Photographed 20 January 2019. Coll. No. 1993, 24 Jul 2018, characters observed: Perennial grass, 30 cm., clumped; Stem, nodes and internodes hidden by leaf sheaths; Leaves, blade and sheath differentiated; Sheath, >50 mm., closed, retrorse sparse long soft hairy, older sheaths persistent at base; Ligule, membranous, 0.5 mm., auricles, absent; Blade, 82 mm. × 3.6 mm. wide, margin, scabrous; Inflorescence, 1 per culm, panicle, 60-70 mm., 1-3 branches per node; Spikelets, #10-25, similar, borne singly, 12-20 mm. × 3-5 mm. wide; Compression, lateral; Disarticulation, above the glumes, between florets; Glumes, lower, 5.8 mm., veins, #1, upper, 8.5 mm., veins, #3, ±parallel (definitely not strongly convergent); Florets, 6 per spikelet, at least lower bisexual, uppermost florets appear sterile with last floret wrapped inside lemma of penultimate floret; Axis, short hairy abaxially; Lemma, 9.5 mm. >glumes, lanceolate, membranous, short ascending soft hairy, thinning distally, awns, #1, 2.3-3.0 mm., attached near tip; Palea, 7.2 mm., <lemma, membranous, tan-brown, veins, green, ciliate; Stamens, #3. Anthers, 2.8 mm.; Achene, 4.5 mm., green, flattened.
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Coll. No. 2638, Bromus ciliatus | Photographed 22 January 2022.
Coll. No. 2638, 5 Aug 2021, characters observed: Perennial grass, 85-90 cm., caespitose; Sheath, closed, basal sheath retrorsely pilose; Ligule, membranous, 1 mm., auricles 0; Blade, 200 mm. × 4 mm. wide, flat, barely scabrous; Inflorescence, 1 per culm, >leaves, open panicle, 90 mm. × 25 mm. wide, 1-3 branches; Rachis, not extended, scabrous; Spikelets, 10-14, 20 mm. × 6 mm. wide, similar, borne singly, pedicels 20-25 mm.; Glumes, lower, 8 mm., veins, 1, somewhat keeled, upper, 9 mm., <lowest floret, veins, 3, keeled, sparsely pilose toward left margin, awns, 0; Florets, 6 per spikelet, #1 is bisexual; Callus, hairs, 0; Lemma, 11.5 mm., body > glumes, lanceolate, membranous, not strongly keeled, veins, 5, tip, entire, awns, 1, 3.8 mm., straight, attached, end, pilose, hairs concentrated proximally and towards the margins, true for upper lemmas; Palea, 10.5 mm., membranous, veins, green, ciliate; Anthers, 1.3 mm.; Achene, 7-8 mm., green, adhering to lemma.
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| Bromus diandrus
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Coll. No. 826, Bromus diandrus | Scanned 8 June 2012.
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| Bromus inermis
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Coll. No. 2843, Bromus inermis | Photographed 10 November 2022.
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Coll. No. 1113, Bromus inermis, auricles absent, or very much reduced. | Scanned 7 September 2015.
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Habitat of Coll. No. 1219, Bromus inermis | Photographed 17 July 2015.
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Coll. No. 1660, Bromus inermis | Photographed 12 December 2017.
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Inflorescence of Coll. No. 1113, Bromus inermis | Scanned 8 September 2015. As the inflorescence matures, it will open somewhat, and become lighter colored.
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Coll. No. 1113, Bromus inermis, characteristic fold on leaf blade. | Scanned 7 September 2015.
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| Bromus japonicus
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Coll. No. 1207, Bromus japonicus | Scanned 6 December 2015. Coll. No. 1207, 13 Jul 2015, characters observed: Annual grass, to 40 cm.; Leaves, sheath, 60 mm. closed + 12 mm. open, pilose (but not densely so), ligule, membraneous, + some hairs, blade, hairy; Inflorescence, panicle, >leaves; Pedicel, 20 mm.; Spikelets, 1-3 per panicle branch, 22 mm. × 6 mm. wide, elliptical, drooping; Glumes, #2, <lowest floret, lower, 4.5 mm., veins 3, upper, 5.5 mm., veins 5; Rachilla, short hairy; Callus, glabrous; Florets, #12, 2-ranked, closely held hiding rachilla; Disarticulation between florets; Lemma, 8 mm. × 4 mm. wide, margin, hyaline, 0.5 mm., tip bifid, awn, #1, 5-12 mm. long, mid-spikelet awns longest, flattened at base, not twisted, attached near bifid tip.
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Coll. No. 1461, Bromus japonicus | Photographed 20 December 2016.
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Coll. No. 1207, Bromus japonicus | Photographed 13 July 2015.
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Coll. No. 1207, Bromus japonicus | Scanned 6 December 2015. Coll. No. 1207, 13 Jul 2015, characters observed: Annual grass, to 40 cm.; Leaves, sheath, 60 mm. closed + 12 mm. open, pilose (but not densely so), ligule, membraneous, + some hairs, blade, hairy; Inflorescence, panicle, >leaves; Pedicel, 20 mm.; Spikelets, 1-3 per panicle branch, 22 mm. × 6 mm. wide, elliptical, drooping; Glumes, #2, <lowest floret, lower, 4.5 mm., veins 3, upper, 5.5 mm., veins 5; Rachilla, short hairy; Callus, glabrous; Florets, #12, 2-ranked, closely held hiding rachilla; Disarticulation between florets; Lemma, 8 mm. × 4 mm. wide, margin, hyaline, 0.5 mm., tip bifid, awn, #1, 5-12 mm. long, mid-spikelet awns longest, flattened at base, not twisted, attached near bifid tip.
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| Bromus lanatipes
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Dissected spikelet of Coll. No. 2170, Bromus lanatipes | Scanned 22 July 2019. Coll. No. 2170, 21 July 2019, characters observed: Perennial grass (short-lived?), 50 cm., clumped; Roots, fibrous; Stem, herbaceous; Internodes, glabrous; Leaves, blade and sheath differentiated, soft; Sheath, 60-70 mm., closed, long shaggy hair; Ligule, 1.5 mm., membraneous, auricles, absent; Blade, 160 mm. × 5 mm. wide, adaxial scaberulous, tip, acute; Inflorescence, 1 per culm, >leaves, panicle, 130 mm. × 60 mm. wide, 2+ branches; Rachis, not extended, scabrous; Spikelets, #12-13, 23 mm. × 5 mm. wide, similar, borne singly; Compression, lateral; Disarticulation, unknown, should be above the glumes; Glumes, lower, 5 mm., veins, #1, keeled, keel scabrous, upper, 6 mm., <lowest floret, veins, #5, keeled; Lemma, 7.5 mm., >glumes, hairy laterally becoming more broadly hairy distally, veins, #5, convergent, awns, #1, 3 mm., straight, barbellate, attached near end; Palea, 9 mm., ±=lemma, membraneous, keel ciliate, veins, lt. green, margin, glabrous, tip, entire; Anthers, #3, 2.3 mm., yellow; Achene, immature.
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Coll. No. 2170, Bromus lanatipes | Photographed 23 January 2020.
Coll. No. 2170, 21 July 2019, characters observed: Perennial grass (short-lived?), 50 cm., clumped; Roots, fibrous; Stem, herbaceous; Internodes, glabrous; Leaves, blade and sheath differentiated, soft; Sheath, 60-70 mm., closed, long shaggy hair; Ligule, 1.5 mm., membraneous, auricles, absent; Blade, 160 mm. × 5 mm. wide, adaxial scaberulous, tip, acute; Inflorescence, 1 per culm, >leaves, panicle, 130 mm. × 60 mm. wide, 2+ branches; Rachis, not extended, scabrous; Spikelets, #12-13, 23 mm. × 5 mm. wide, similar, borne singly; Compression, lateral; Disarticulation, unknown, should be above the glumes; Glumes, lower, 5 mm., veins, #1, keeled, keel scabrous, upper, 6 mm., <lowest floret, veins, #5, keeled; Lemma, 7.5 mm., >glumes, hairy laterally becoming more broadly hairy distally, veins, #5, convergent, awns, #1, 3 mm., straight, barbellate, attached near end; Palea, 9 mm., ±=lemma, membraneous, keel ciliate, veins, lt. green, margin, glabrous, tip, entire; Anthers, #3, 2.3 mm., yellow; Achene, immature.
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| Bromus madritensis rubens
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Coll. No. 1589, Bromus madritensis ssp. rubens | Scanned 19 November 2017. Coll. No. 1589, 18 April 2017, characters observed: Annual grass, to 25 cm., soft hairy throughout; Sheath (... looks open to me ...) Ligule, membraneous, 4 mm., laciniate; Inflorescence, panicle; Compression, lateral; Spikelets, 1 per node, 27 mm., axis mostly glabrous; Glumes, lower, 7 mm., upper, 9 mm., veins 5, keeled (IMHO); Florets, #4-6; Lemma, 13 mm., veins 3, tip bifid, awn, #1, 9 mm., attached to back; Palea, 9 mm., margin ciliate; Grain, 8 mm., flattened, brown.
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Coll. No. 1570.2, Bromus madritensis ssp. rubens | Photographed 3 October 2017. Coll. No. 1570.2, 18 April 2017, charcters observed: Annual grass, to 20 cm.; Inflorescence, 50 mm.; Spikelets, 40 mm., similar; Compression, lateral (looks strongly lateral to me, but this will send you down the wrong path); Glumes, lower, 7 mm., slightly keeled, few hairs, veins, 1, upper, 10 mm., rounded, veins, 3; Florets, 6+; Lemma, 13 mm. × 2.3 mm. wide, veins, 3, tip, bifid, awns, 1, 11 mm., attached to back; Palea, 9 mm., margin sparsely ciliate; Grain, 9 mm. × 1.3 mm. wide, flattened.
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| Bromus porteri
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Coll. No. 2860, Bromus porteri. | Photographed 21 November 2022.
Coll. No. 2860, 18 July 2022, characters observed: Perennial grass, 45 cm., caespitose; Leaves, sheath, 130 mm., open about ½-way; collar minutely hairy, ligule, 2 mm., blade, 130 mm. × 4 mm. wide, flat, scabrous; Inflorescence, nodding, 1 per culm, panicle; Rachis, ascending hairy, not quite appressed; Spikelets, many, 23 mm. × 2.8 mm. wide, similar, pedicels, 44 mm.; Compression, unremarkable; Glumes, soft spreading hairy, lower, 7.5 mm., veins, 3, upper 8.5 mm., < lowest floret, veins, 3, converging, awns, 0, but mid-vein goes strongly right to the tip; Florets, 8 per spikelet; Callus, few short hairs; Lemma, 10.5 mm., > glumes, rolled, hairy. veins, 5, convergent, awns, 1, 1 mm., subterminal, straight; Palea, 9.5 mm., < lemma, membranous, ciliate, tip rounded; Stamens, 3; Anthers, 2.2 mm.
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| Bromus pumpellianus
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Obscure rhizome on Coll. No. 2176, Bromus pumpellianus | Scanned 27 January 2020.
Coll. No. 2176, 23 Jul 2019, characters observed: Perennial grass, to 70-80 cm., occasionally obscurely rhizomatous; Roots, fibrous; Stem, herbaceous; Leaves, blade and sheath differentiated, soft, sheath, 90 mm., closed, ligule, membraneous, 1 mm., auricles, minimal, blade, 90 mm. × 3 mm. wide, flat, scabrous, margins, scabrous; Inflorescence, 1 per culm, >leaves, panicle, 130 mm. × 30 mm. wide; Rachis, not extended, scabrous; Spikelet, many, 26 mm. × 5 mm. wide, similar, 1 per node, pedicels 16-30 mm.; Compression, lateral; Disarticulation, above the glumes, between florets; Glumes, lower, 6 mm., veins, ≥3, back hairy, upper, 8.5 mm., <lowest floret, veins, ≥3, rounded, membraneous, awns, 0, back hairy, somewhat more dense distally; Florets, 6 per spikelet, bisexual; Lemma, 10 mm., >glumes, lanceolate, rolled, uniformly hairy on back, green, veins, #7, convergent, tip, rounded, awns, #1, 1.5 mm., straight, attached back of lemma, near tip; Palea, 8 mm., <lemma, membraneous, back hairy, keel ciliate, veins, green, margin, glabrous, tip, entire; Anthers, 2.3 mm.; Ovary, glabrous; Stigma, feathery; Achene, 3 mm., green.
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| Bromus secalinus
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Coll. No. 2362, Bromus secalinus | Photographed 12 December 2020.
Coll. No. 2362, 20 June 2020, characters observed: Annual grass, to 60 cm.; Leaves, sheath, 85 mm., closed 70 mm., glabrous, ligule, membraneous, erose, blade, 100 mm., adaxially hairy; Inflorescence, open panicle; Spikelets, 17 mm. × 5 mm. wide (excl. awns); Glumes, <lowest floret, lower, 5 mm., veins, #5, upper, 7 mm., veins, #7; Disarticulation, between florets; Callus, glabrous; Lemma, 7.5 mm. × 3.5 mm. wide (broad), rolled (not keeled), veins, #7, converging; Awn, 6 mm., straight, attached back, 1.3 mm. fom apex; Fruit, U-shaped.
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| Bromus sterilis
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Coll. No. 2841, Bromus sterilis. | Photographed 8 November 2022.
Coll. No. 2841, 2 Jul 2022, characters observed: Annual grass, 22-33 cm.; Leaves, sheath, closed, scantily pilose, ligule, membranous, 1.5 mm., lacerate, auricles, absent, blade, 15 cm., × 1.8 mm. wide, flat; Inflorescence, 1 per culm, >leaves, panicle, 14 cm. × 2.5 cm. wide; Rachis, not extended, scabrous; Spikelets, #5, 42 mm. × 5 mm. wide, similar, borne singly, pedicels, 25 mm., Compression, lateral; Glumes, lower, 9 mm., upper 13.5 mm., <lowest floret; Florets, 6 per spikelet; Axis, scabrous; Callus, minutely hairy; Lemma, 16.5 mm., >glumes, oddly rolled rather than keeled, mid-vein to margin 0.9-1.1 mm., awn, subterminal, 17.5 mm., straight.
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| Bromus tectorum
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Coll. No. 1112, Bromus tectorum | Scanned 6 September 2015. Coll. No. 1112, 26 May 2015, characters observed: Annual grass, to 30 cm.; Leaves, Sheath, 50 mm., closed proximal ⅔, open distally, Ligule, membraneous, 1 mm.; Blade, 60 mm.; Inflorescence, open panicle, rachis and pedicels short hairy, Spikelets, 1 per node, 15 mm., laterally compressed; Glumes, 2, sparsely hairy, margins scarious, lower, 2.3 mm., veins indistinct, upper, 4.5 mm. Callus, glabrous, very short, Florets, ±6, Lemma, 6.5 mm., 0.7 from mid-vein to margin, back rounded, sparsely hairy, tip, forked, teeth 1 mm., awned from near tip, awn, 5-7 mm.; Anthers, 3, 0.4 mm. Dimensions are small for B. tectorum, perhaps because of the early date of collection.
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Coll. No. 825, Bromus tectorum | Scanned 16 Jun 2013. I had keyed this out at B. carinatus, but it has since been annotated as B. tectorum. Coll. No. 825, 30 May 2012, characters observed while keying: Perennial, to 40 cm.; Roots fibrous; Stem herbaceous; Internodes hollow; Leaf blade and sheath differentiated; Sheath closed; Ligule membraneous, 2 mm.; Blade flat, to 4 mm wide, short hairy, margin sparse wavy hairy; Inflorescence 3-many, 10-12 cm., panicle, branching evident, longest branch 25 mm., > leaves, spikelets many; Compression lateral; Spikelets similar, 1 / node, 1° and 2° branches, length 30-35 mm., 5-6 florets; Florets 2+ bisexual; Glumes lower 7 mm., upper 10 mm., margins thin translucent, upper glume has 3 prominent veins (keeled), awns 0; Lemma, lower 12 mm., keeled, veins = 3 or 7, outer 2 appear doubled, tip 2-forked, awns 1, 13 mm, straight, attached to back at 9 mm.; Palea, 8 mm, < lemma, keel hairy, margin ciliate; Anthers 3, 0.7 mm., brown; Ovary 5 mm., glabrous below, hairy near tip.
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| Buchloe dactyloides
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Pistillate inflorescence of Buchloe dactyloides | Scanned 12 August 2018.
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Pistillate inflorescence of Buchloe dactyloides | Scanned 12 August 2018.
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Pistillate inflorescence of Buchloe dactyloides | Scanned 12 August 2018.
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Coll. No. 1673, Buchloe dactyloides | Scanned 16 December 2017.
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Staminate inflorescence of Coll. No. 1673, Buchloe dactyloides | Scanned 16 December 2017.
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| Calamagrostis canadensis
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Coll. No. 2700, Calamagrostis canadensis. | Photographed 9 February 2022.
Coll. No. 2700, 17 August 2021, characters observed: Perennial grass, to 90 cm., rhizomes unknown; Leaves, sheath, 90 mm., open, ligule, membranous, 3-5 mm., collar, minutely hairy, blade, 150 mm. × 6.5 mm. wide, scabrous; Inflorescence, 1 per culm, >leaves, 135 mm. × 25 mm. wide; Spikelets, many, 3 mm. × 1 mm. wide, similar, borne singly; Compression, ±lateral; Disarticulation, above the glumes; Glumes, lower, 3.5 mm., veins, 1, scabrous, purple to brown, upper, 3.0 mm., >lowest floret, scabrous, veins, 3, rounded, purple to brown; Florets, 1 per spikelet; Callus, straight white hairs, 2.5 mm.; Lemma, 2.5-3.0 mm., body < glumes, tip, 2-forked, awns, 1, 2.5 mm., straight, attached middle, included; Palea, 2 mm., <lemma, membranous. Compare this collection to my Coll. No. 2011 (KHD, OBI, UCR), which was collected in earlier in the season of 2018 about 700 m. southeast of this collection. It was determined C. stricta at the time, but is very similar to this one. Perhaps, it is also C. canadensis. Per SEINet, there are six collections of Calamagrostis within 15 km. of my two collections; four of C. canadensis and two of C. stricta.
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| Calamagrostis purpurascens
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Coll. No. 3041, Calamagrostis purpurascens. | Photographed 21 December 2023.
Coll. No. 3041, 3 July 2023, characters observed: Perennial grass, to 40 cm., no rhizomes seen; Leaves, sheath, 12 cm., open, ligule, membranous, 4 mm., auricles, 0, blade, 15 cm. × 3 mm. wide, flat, scabrous, also adaxially densely and minutely hairy between the ribs; Inflorescence, <leaves, 1 per culm, panicle, 110 mm. × 12 mm. wide; Rachis, not extended; Spikelets, many, 8 mm. × 1 mm. wide, similar; Glumes, lower, 6 mm., veins, 1, upper, 5 mm., > lowest floret, veins, 1 (3?), keeled, awns, 0; Florets, 1* per spikelet; *Rachilla, extended beyond upper floret as though there could be another floret up there, hairy; Callus, short straight white hairs; Lemma, 4 mm., < glumes, green, veins, 5, glabrous, tip, toothed, awns, 1, 7 mm., bent, attached, near base of lemma; Palea, 4 mm., < lemma, membranous; Achene, 1.7 mm., green.
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| Calamagrostis stricta
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Coll. No. 2011, Calamagrostis stricta | Photographed 27 January 2019. Coll. No. 2011, 25 July 2018, characters observed: Perennial grass, 90-100 cm., rhizomatous; Leaves, sheath, 120 mm., open, minutely scabrous; ligule, membranous, 4 mm., auricles, absent, collar, minutely hairy; blade, 160 mm. × 3.2 mm. wide, scabrous; Inflorescence, 1 per culm, >leaves, contracted panicle, 150 mm. × 15 mm. wide; Spikelets, many, 3.8 mm. × 0.8 mm. wide, similar, 1 per node; Compression, ±lateral; Disarticulation, above the glumes; Glumes, lower, 3.8 mm., veins, #1, upper, 3.2 mm., >lowest floret, veins, #3, keeled proximally, keels scabrous, awns, #0; Florets, 1 per spikelet; Callus, straight white hairs, 2.5 mm., ≤lemma; Lemma, 3 mm., <glumes, tip, 2-forked; awns, #1, 2.5 mm., straight, attached below middle; Palea, 1.9 mm., <lemma, membranous.
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| Calamovilfa longifolia
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Spikelets of Coll. No. 2397, Calamovilfa longifolia | Scanned 23 December 2020.
Coll. No. 2397, 29 Jul 2020, characters observed: Perennial grass, 1 m., thick scaly rhizomes; Nodes, hidden by sheath; Leaves, blade and sheath differentiated; Sheath, >75 mm., open, glabrous to long straight hairs distally; Ligule, hairs, 1.5 mm.; Auricles, absent; Blade, 350 mm. × 7 mm. wide, flat; Inflorescence, 1 per culm, panicle, 180 mm. × 15 mm. wide, 3-4 branches; Rachis, glabrous, not extended; Spikelets, many, 5.5 mm. × 2.5 mm. wide, borne singly, subtended by few long silky hairs; Compression, lateral. Disarticulation, above the glumes; Glumes, lower, 3.5 mm., keeled, veins, #1, upper, 5 mm., < lowest floret, veins, #1, keeled, awns, absent; Callus, straight silky hairs, 3 mm.; Lemma, 5 mm., >glumes, lanceolate, keeled, glabrous, scarious, veins, #1, forming a tiny awn tip, awns, absent; Palea, 4.8 mm., <lemma, colorless, glabrous; Stamens, #2; Anthers, 3.8 mm.
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Florets of Coll. No. 2397, Calamovilfa longifolia | Long silky hairs on the callus. Scanned 23 December 2020.
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