Eastern Mojave Vegetation
| Detailed Notes About Colorado Taxa (Continued) |
|
Edited by Tom Schweich |
Topics in this Article: Appendices Notes on Native and Non-Native Plants Ferns and Allies Gymnosperms — Conifers and Allies Dicots Monocots Literature Cited |
Unified set of notes regarding plants. Pages:
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Dicots | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Populus L. “Poplar” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Populus × acuminata Rydb. “Lanceleaf Cottonwood” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Literature Cited:
Locations: Carter Canyon. |
Rydberg, Per Axel 1893. On the American Black Cottonwoods. Bulletin of the Torrey Botanical Club. 20(2) pp.46-50 https://www.biodiversitylibrary.org/item/7998#page/72/
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Populus alba L. “White Cottonwood” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Populus angustifolia E. James “Narrowleaf Cottonwood” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Literature Cited:
Locations: Brighton. |
James (1822, v. 1, p. 497) ... entry for July 4, 1820, while encamped on the South Platte River near Brighton.
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Literature Cited:
Locations: Woodland Park. |
Rydberg, Per Axel 1893. On the American Black Cottonwoods. Bulletin of the Torrey Botanical Club. 20(2) pp.46-50 https://www.biodiversitylibrary.org/item/7998#page/72/
Manitou is a variant name for Woodland Park. GNIS has no entries for War Bonnet Canyon or Little Elk Canyon. Although, there are three Little Elk Creeks, a Little Elk Basin, and a Little Elk Park in Colorado.
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Populus deltoides Marshall ssp. monilifera (Aiton) Eckenw. “Plains Cottonwood”(Syn: Populus sargentii Dode) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Literature Cited:
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Marshall (1785, p. 106) ...
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Literature Cited:
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Aiton (1789, v. 3, p. 406) ...
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Literature Cited:
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Eckenwalder (1977, v. 58, n. 3, p. 93-208) ...
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Literature Cited:
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“ ... [like a string of beads, alluding to the racemes of fruits] ...” (Weber & Wittmann, 2012, p. 347). | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Populus tremuloides Michx. “Quaking Aspen” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Literature Cited:
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Michaux (1803, v. 2, p. 243) ...
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Salix alba L. “White Willow” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Linnaeus (1753, v. 2, p. 1021) ...
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Salix amygdaloides Andersson “Peachleaf Willow” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Literature Cited:
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Andersson, N. J. 1858. Enumeratio Salicum nobis ex America boreali huc usque cognotarum “The Willow Flora known to us from north America” Öfversigt af Kongl. Vetenskaps-akademiens forhandlingar “Overview of the Proceedings by the Royal Academy of Sciences” v. 15, p. 114-133 https://www.biodiversitylibrary.org/item/54173 Andersson (1858, v. 15, p. 114) …
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Salix exigua Nutt. “Coyote Willow” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Literature Cited:
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The species was described by Nuttall (1842, vol. 1, p. 90) in his extension of Michaux's North American Sylva as being found in the Territory of Oregon without any additional information about collector, location, date, &c. At the time Oregon extended from central Wyoming to the Pacific coast.
It is a little curious that Nuttall mentions no willows in either his report of his residency in Oregon (Nuttall, 1840) or in his descriptions of plants collected by William Gambel (Nuttall, 1848). | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Literature Cited:
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The treatment of Salix in FNANM, written by Geoge Argus, treats S. exigua with one variety hindsiana, which is not known to occur in Colorado. Argus (2010) treats S. interior at the rank of species saying:
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Weber & Wittmann (2012) do not accept any varieties of S. exigua, treating S. interior at the rank of species. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Literature Cited:
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Ackerfield (2015) treats S. interior Rowlee as S. exigua var. interior (Rowlee) Cronquist. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Salix irrorata Andersson “Dewystem Willow” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Literature Cited:
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Andersson (1858, v. 15, p. 117) ...
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Juglans nigra L. “Black Walnut” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Linnaeus. 1753. vol. 2, p. 997
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Alnus incana (L.) Moench ssp. tenuifolia (Nutt.) Brietung “Thinleaf Alder” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Betula occidentalis Hook. “Water Birch” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Hooker. [1829-1840.] 1838, v. 2, pt. 10, p. 155. Flora boreali-americana.
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Carpinus betulus L. “European hornbeam”There is only one collection of Carpinus betulus L. “European hornbeam” in Colorado, that made near the Coors plant in Golden in 1949. Linnaeus (1753) described the tree as native to Europe and Canada, though POWO does not show it lives there, nor are there any Canadian collections in SEINet. The tree is native to Germany, and it is possibly planted on the Coors grounds for that reason. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Linnaeus (1753, v. 2, p. 998) ...
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Corylus cornuta Marshall “Beaked Hazelnut” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Quercus gambelii Nutt. “Gambel Oak”Gambel Oak — Quercus gambelii — is found mostly in southern Jefferson County and, until recently, not in Golden. However, the writer found it recently in a canyon in Apex Park. It is also found on Dinosaur Ridge just south of Interstate 70 and therefore just south of the Golden city limits. The Gambel oaks on Eagle Ridge Drive near Kinney Run were planted. The oak is broadly distributed throughout the American Southwest. For example, the writer has also collected it in the Spring Mountains, Clark County, Nevada, about 45 km. northwest of Las Vegas.
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Literature Cited:
Locations: Rio Grande. |
William Gambel (June 1823 – December 13, 1849) was an American naturalist, ornithologist, and botanist from Philadelphia. As a young man he worked closely with the renowned naturalist Thomas Nuttall. At the age of eighteen he traveled overland to California, becoming the first botanist to collect specimens in Santa Fe, New Mexico and parts of California. In March 1841, at the age of eighteen, Gambel set off on his own for California to collect plants and other specimens for Nuttall. He planned to take a more southerly route than that taken in 1834 by Nuttall and John Kirk Townsend. Upon reaching Independence, Missouri he joined a group of traders and headed for Santa Fe following the Santa Fe Trail. Gambel reached Santa Fe in June and spent the next couple months collecting plants. In September Gambel joined a party heading to California and accompanied them along the Old Spanish Trail, arriving in Mexican Alta California in early November, 1841, becoming the first botanist to enter California overland from the east. Nuttall (1848) described an oak collected on the Rio Grande by William Gambel.
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The collections around Golden may be the northernmost occurrences along the Front Range. Gambel oak has been collected in the vicinity of Boulder, though Weber & Wittmann (2012) note that they were planted. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Ulmus americana L. “American Elm” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Ulmus parvifolia Jacq. “Chinese Elm” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Ulmus pumila L. “Siberian Elm” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Other articles:
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Celtis occidentalis L. “Western Hackberry” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Literature Cited:
Other articles:
|
| ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Other articles:
|
Celtis reticulata Torr. “Net-Leaved Hackberry” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Literature Cited:
|
Torrey (1828, v. 2, p. 247) in his description of plants collected by Dr. Edwin James ...
| ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
|
Notes on Humulus L. “Hops” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Literature Cited:
|
Linnaeus (1753, v. 2, p. 1028) ...
| ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Literature Cited:
|
Nuttall (1848, p. 23) proposed Humulus americanus in describing some collections by Gambel.
| ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Literature Cited:
|
Nelson & Cockerell (1903, v. 16, p. 45) ...
| ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Literature Cited:
|
Rydberg (1917) Flora of the Rocky Mountains …
| ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Literature Cited:
|
Harrington (1964, 2nd edition) ...
| ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Literature Cited:
|
Small (1978, ____) published H lupulus var. lupuloides. A numerical and nomenclatural analysis of morpho-geographic taxa of Humulus. — behind a paywall. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Literature Cited:
|
The relationships of hop cultivars and wild variants of Humulus lupulus | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Literature Cited:
|
Löve (1982, p. ___) published H. lupulus subsp. americanus (Nutt.) Á.Löve & D.Löve. — behind a paywall. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Literature Cited:
|
Key in FNANM
| ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Literature Cited:
|
Molecular phylogeny of wild Hops, Humulus lupulus L. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
|
Literature Cited:
|
Weber & Wittmann (2012) refer our local “hops” to Humulus lupulus var. neomexicanus Nelson & Cockerell.
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Literature Cited:
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Ackerfield, 2015, pp. 263 ...
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Literature Cited:
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Tembrock, et al., 2016, p. 19 ...
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Literature Cited:
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Ackerfield, 2022, pp. 282-283 ...
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Literature Cited:
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Humulus lupulus L. “Wild Hops”See the section above “Notes on Humulus sp.” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Other articles:
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Humulus neomexicana (A. Nelson & Cockerell) Rydb. “New Mexican Hops”See the section above “Notes on Humulus sp.” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Other articles:
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Parietaria pensylvanica Willd. “Pennsylvania Pellitory” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Other articles:
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Urtica dioica L. ssp. gracilis (Aiton) Seland. “Stinging Nettle” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Other articles:
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Arceuthobium vaginatum (Willd.) J. Presl var. cryptopodium (Engelm.) Cronquist “Pineland Dwarf Mistletoe” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Literature Cited:
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Gray (1850, v. 6, p. 214) includes a long communication from George Engelmann regarding the mistletoes.
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Other articles:
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Comandra umbellata (L.) Nutt. ssp. pallida (A. DC.) Piehl. “Pale Bastard Toadflax” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Literature Cited:
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Linnaeus (1753) first described Comandra umbellata from plants collected in Virginia and Pennsylvania.
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Literature Cited:
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Nuttall (1818) defined a new genus, Comandra, and places the North American Thesium into it. Nuttall also suggests that some of Linnaeus' eastern hemisphere Thesium might belong in Comandra. The genus Thesium L. remains accepted, and is native to Europe, Asia-Tropical, Africa, South America and Asia-Temperate.
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Literature Cited:
Other articles:
Locations: Lapwai. |
DeCandolle (1857) proposed C. pallida from a collection in Oregon Territory.
Another first collection by Spalding was Nothocalais troximonoides. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Oliphant (1934) published a paper on the botanical labors of the Reverend Henry H. Spalding. Spalding collected for one year and sent one box of dried plants east. He was apparently stimulated to this activity by contact with Charles A. Geyer, a German botanist, who was in Oregon Country in 1843-44. Spalding sent one box in 1846, which was received in Boston in 1848. By the time that Gray's encouragement for Spalding to continue collecting was sent to him in March 13, 1849, a massacre at Waiilatpu had sent Spalding and others fleeing to Lower Oregon ending his botanical career. I don't know how Spalding's collections ended up in DeCandolle's herbarium, and why it was he and not Asa Gray who published Spalding's Comandra collections. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Piehl (1965) wrote a natural history and taxonomy of Comandra. It is almost a monograph, but I don't think we do monographs about single species.
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Der and Nickrent (2008) … phylogenetic analysis supports a Comandra clade consisting of Comandra and Geocaulon lividum, i.e., the C. livide rejected by DeCandolle. Their work also supports a Thesium clade distinct from a Comandra clade. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Two hemi-parasites were encountered in species sampling: Comandra umbellata, and Pedicularis canadensis. C. umbellata was a significant indicator for high-quality prairies. (Sivicek & Taft, 2011) | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Eriogonum Michx. “Hairy Plant” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Michaux (1803, Vol. 1, pg. 246) published the first Eriogonum meaning “hairy plant.”
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Subgroups of Eriogonum as described by Reveal (2005) in his treatment of the genus in Flora of North America.
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Those that have been collected in Golden are found in three of the subgroups:
Two other wild buckwheats that are often of interest are both in subgroup Oligogonum Nutt.:
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Eriogonum Michx. — Wild Buckwheat1a. Perennials, low, spreading, cespitose or mat-forming; flowers attenuate at base, sometimes only weakly so, stipelike base not at all winged; inflorescence bracts 2-10 or sometimes more; perianths glabrous or, if pubescent, not white-tomentose ... 3. (Eriogonum subg. Oligogonum) 1b. Perennial subshrubs or tall erect herbs; flowers not attenuate at base, stipelike base absent or, if present, then slightly winged; inflorescence bracts usually 3 ... 2 2a. Plants perennial subshrubs; involucres (except 1 at fork of node in some) typically sessile, without winged or distinctly ridged achenes ... 6. (Eriogonum subg. Eucycla) 2b. Tall erect perennial herb, Involucres pedunculate, flowers usually glabrous, achenes winged or distinctly ridged ... Eriogonum subg. Pterogonum. Achenes 3-winged; plants (3) 5-25 dm. tall, from a thick taproot; leaves all or mostly basal, linear-lanceolate to oblanceolate, hirsute, 5-20 cm long; flowers yellow of yellowish green, or reddish in fruit … E. alatum 3. (Eriogonum subg. Oligogonum) 3a. Perianth glabrous externally; stems only with a whorl of leaf-like bracts subtending the inflorescence; ... E. umbellatum 3b. Perianth hairy externally, stems with a whorl of leaf-like bracts subtending the inflorescence and reduced or scale-like bracts at distal nodes; … 4 4a. Flowers white, cream, or pinkish … E. jamesii var. jamesii (in Jeffco known only from an M. E. Jones' collection in Morrison, 1878). 4b. Flowers pale to bright yellow … 5 5a. Involucres elevated on evident peduncles … E. flavum var. flavum 5b. Involucres sessile or nearly so … E. arcuatum 6. (Eriogonum subg. Eucycla). Leaves 4-9 times longer than wide (to 1 cm wide), linear-lanceolate, narrowly oblong, narrowly ellliptic, or oblanceolate, the margins not crenulate. Plants of the eastern slope … E. effusum | |||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Eriogonum subgroup Eucycla Nutt. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Nuttall (1848b, p. 166) published Eucycla as a genus name for wild buckwheats he had collected, probably on his 1834 expedition to Oregon Territory.
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The characters of subgroup Eucycla are primarily: perennial, stems not jointed, flowers without a stipe-like base. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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The one representative of subgroup Eucycla that is found in Golden s.l. is E. effusum Nutt. “Spreading Buckwheat.” Other wild buckwheats in subgroup Eucycla in the author's experience are E. microthecum Nutt. “Slender Buckwheat” from the Mono Lake Basin and E. wrightii Torr. ex Benth. “Bastardsage” from the eastern Mojave Desert. All three of these wild buckwheats are very similar appearing. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Eriogonum subgroup Oligogonum Nutt.Nuttall (1848b, p. 165) discusses the differences between E. tomentosum on which Michaux (1803) founded the genus Eriogonum, noting some key differences and proposing a subgenus Oligogonum which contains some of our more common wild Buckwheats.
Oligogonum is the larger of the subgroups and contains many of the caespitose or mat-forming perennials, those with a stipe-like base on the flowers. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Eriogonum subgroup Pterogonum GrossGross (1913, vol. 49, pg. 239, footnote 1) in his “Contributions to the knowledge of the Polygonaceae” proposed Pterogonum for some Eriogoneae but I cannot read the original German well enough to understand the basis for segregation from the remainder of Eriogoneae.
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Reveal (2005) places six North American taxa in subgroup Pterogonum of which only one, E. alatum occurs in Colorado. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Eriogonum alatum Torr. “Winged Buckwheat” | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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No mention of Eriogonum alatum in the report of Frémont's 1842 and 1843-'44 expeditions, though Torrey (1853) notes they were aware of it. | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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No mention of Eriogonum alatum in Gray's (1852) account of Wright's collections in Texas and New Mexico in 1849. In fact, there are no Polygonaceae in either part of Gray's account, although DeCandolle (1857) will say he has seen a collection of Eriogonum alatum by Wright. Gray's preface notes that the cacti were given to George Engelmann, of St. Louis, for examination. The seeds were divided between the Botanic Garden of Harvard University, under Gray's charge, and the Royal Botanic Gardens at Kew, under the direction of Sir Wm. Hooker. Sets of plants were distributed to Gray's herbarium, the Smithsonian Institution, John A. Lowell, a patron of the Wright's explorations, with a few distributions to others. If one assumes that Wright collected any Polygonaceae, what happened to them? | ||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||||
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Dr. John Torrey wrote the Botany chapter of the Sitgreaves Report on the expedition Sitgreaves led down the Zuni and Colorado Rivers in 1851.
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• Golden Checklist Flora:
Solanum triflorum;
Literature Cited:
- Nuttall, Thomas, 1818.
Other articles:
• Publication Details:
Nuttall, 1818, publication details;
Nuttall (1818) described the Solanum of North America and included a review of Solanum and related taxa, their origins and uses.
Original Text Comments 6. *triflorum. Stem unarmed, herbaceous and procumbent; leaves dentately-pinnatifid, smooth, segments acute, somewhat undulated, with the margin more or less revolute; peduncles opposite the leaves, 2 or 3-flowered. — Flowers small and white, revolute; fruit about the size of a cherry, green when ripe. Stem a little hirsute, spreading and procumbent, about a foot long; leaves somewhat runcinate. This species, though very distinct, appears to have some affinity with the S. runcinatum of Peru and Chili (sic). — Hab. As a weed in and about the gardens of the Mandans and Minitarees, and in no other situations. Near Fort Mandan. Flowering from June to August. This is followed by a several paragraph review of Solanum and related species, their origins and uses.
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• Golden Checklist Flora:
Limosella aquatica;
Other articles:
• Golden Checklist Flora:
Scrophularia lanceolata;
Other articles:
• Golden Checklist Flora:
Verbascum blattaria;
Other articles:
• Golden Checklist Flora:
Verbascum phlomoides;
Literature Cited:
- Mitchell, John D., 1769.
Other articles:
• Taxa Notes:
Notes on Orobanche;
Literature Cited:
- Gray, Asa, 1856.
Other articles:
• Taxa Notes:
Notes on Orobanche fasciculatum, Gray, 1856;
Gray, Asa. 1856. 2nd ed. Manual of the botany of the northern United States. New York. Publication of Aphyllon fasciculatum
4. APHYLLON, Mitchell. Naked Broom-rape. Flowers perfect, solitary on long naked scapes or peduncles, without bractlets. Calyx 5-cleft, regular. Corolla with a long curved tube and a spreading border, somewhat 2-lipped; the upper lip deeply 2-cleft, its lobes similar to the 3 of the lower lip. Stamens included. Stigma broadly 2-lipped. Capsule with 4 equidistant placentae, 2 borne on each valve half-way between the midrib and the margin. Plants brownish or yellowish. Flowers purplish, and scapes minutely glandular-pubescent. (Name from α privative and Φυλλον, foliage, alluding to the naked stalks.) — Perhaps rather a section of Phelipaea. ;
Other articles:
• Golden Checklist Flora:
Castilleja integra;
• Field Notes:
Coll. No. 1143, 15 Jun 2015;
Coll. No. 1379, 29 May 2016;
Coll. No. 1614, 15 May 2017;
Coll. No. 1865, 23 May 2018;
A very common paintbrush in Golden s.l., Castilleja integra A. Gray “Wholeleaf Indian Paintbrush” is found in all the open spaces. Jefferson County collections are mainly in the northern part of the county, on the plains and up in the foothills. Most of the Colorado state collections are in the Rocky Mountains and adjacent plains from Denver south.
The name was applied by A. Gray (1859) in “Pacific Railroad Reports” while citing a number of contemporary collections in the southwest.
In 1849, he joined an army expedition (with Gray's help) through Texas, botanising from Galveston to San Antonio and then on to El Paso. But he had to walk most of the 673 miles, (which took over 104 days effort). He collected seeds of Penstemon baccharifolius (Hook.), between Texas and El Paso, which were later given to William Hooker. Also, Castilleja lanata (found near the Rio Grande) and Castilleja integra (found in the Organ Mountains, near El Paso) (Pennell, 1935). In the spring of 1851, he joined the United States and Mexican Boundary Survey (also with Gray's help). (Wikipedia)
Literature Cited:
- Gray, Asa, 1849.
Other articles:
• Publication Details:
Gray, 1849, publication details;
Castilleja integra is not listed in Gray (1849) Plantae Fendlerianae Novi-Mexicanae. Although, a good part of the report is "… to be continued."
Literature Cited:
- Gray, Asa, 1852-1853.
Gray (1952) does not mention any Wright collections in the Scrophulariaceae
Literature Cited:
- Torrey John, 1859.
Emory, William H., 1859. Report on the United States and Mexican Boundary Survey. Volume II. Torrey, John, 1859. Part I. Botany of the Boundary Washington, 1859. p. 119
SCROPHULARIACEAE (by A. Gray)
…
Castilleja integra (sp. nov.): perennis; caule stricto tomentoso; foliis linearibus integerrimis subtus tomentulosis, floralibus oblongis obocatisque integerrimis coloratis (paniceis);; spica conferta; calyce aequaliter vel postice profundius bifido, lobis bifidis lanceolatis obtusiusculis labium inferius galea multoties brevius adaequantibus. — Organ mountains, east of El Paso; Wright, (undistributed,) Bigelow. Guadaloupe cańon, Sonora; Capt. E. K. Smith. Also gathered in the Rocky Mountains further north by Dr. Kreuzfeldt, in Gunnison's expedition. Stem one or two feet high, mostly simple, rigid; leaves 1˝ to 3 inches long, 2 to 3 lines wide, entire; most of the floral ones almost wholly petaloid, ample, shorted than the fully developed flowers. Calyx 8 or 12 lines long, red or reddish; “corolla reddish green;” glaea 6 to 8 lines long; the lower lip very short. Apparently a well marked new species of the section Callichroma. It is No. 584 of Fendler's New Mexican collection; and Dr. Bigelow gathered specimens in Whipple's expedition on the Llano Estacado.
…
Literature Cited:
- Parry, Charles C., 1862.
Other articles:
• Golden Checklist Flora:
Parry, 1861;
• Publication Details:
Parry, 1862b, Publication details;
Castilleja integra was among Parry's 1861 collections. Parry's collection (GH1877286) is probably the first and oldest collection of the taxon from Colorado. Location data is the typical “From the head-waters of Clear Creek, and the alpine ridges lying east of "Middle Park,"
Literature Cited:
- Gray, Asa, 1863.
Other articles:
• Golden Checklist Flora:
Parry, Hall & Harbour, 1862;
Castilleja integra was among the collections of Parry, Hall, and Harbour, but Gray (1863) gives no location data. There are two vouchers from 1862 in herbaria, PBRU44553, and COLO744060, neither with locality data.
Literature Cited:
- Pennell, Francis W., 1935.
Pennell, Francis W. [Curator of Botany, Academy of Natural Sciences of Philadelphia] The Scrophulariaceae of Eastern Temperate North America The Academy of Natural Sciences Monographs, Number 1 Philadelphia, 1935. p. 533
9. Castilleja integra Gray Castilleja integra A. Gray, in Torrey, Bot. Mex. Bound. 119. 1859. “Organ mountains, east of El Paso; Wright …, Bigelow. Guadaloupe canon, Sonora; Capt. E. K. Smith."
Bracts red. Flowering from May to August.
Gravelly soil, plains and hills, Colorado to Texas, Chihuahua and Arizona. Known in our territory from a single record along the Rio grande.
Texas. Valverde: bluffs of Devils R., Havard (U).
Other articles:
• Taxa Notes:
Notes on Aphyllon, Mitchell, 1769;
Several Orobanche species are known to be agricultural weeds in Colorado, and some have been reported to infest agricultural lands. Species of Concern:
- Orobanche cooperi (Cooper's broomrape): While generally not considered a severe weed, it has been found infesting agricultural cropland, including tomato fields in inland California. Although its primary hosts are plants in the Asteraceae family like Artemisia and Ambrosia, it's capable of parasitizing other plants.
- Orobanche fasciculata (Clustered broomrape): This species is commonly found in Northeast Colorado and is known to parasitize sagebrush and other plants. It has also been reported to parasitize various Salt Bushes and other plants in Pinyon/Juniper forests.
- Orobanche minor (Small broomrape): This species is a federally listed noxious weed. There's concern that it may spread to infest crops such as legume forages and leafy green vegetables, and heavy infestations can lead to crop failure.
- Orobanche aegyptiaca (Egyptian broomrape): While a widespread agricultural pest globally, infecting various crops including tomato, eggplant, potato, sunflower, and legumes, the provided search results don't explicitly confirm its presence and impact as a weed in Colorado agriculture.
Literature Cited:
- Piwowarczyk, Renata, et al., 2021.
Piwowarczyk (2021) does not address American Aphyllon in their text. However, their historical biogeography (Figure 3) shows Aphyllon as a strictly American genus diverging roughly 9 million years ago.
Other articles:
• Golden Checklist Flora:
Orobanche fasciculata;
Literature Cited:
- Nuttall, Thomas, 1818.
Described by Nuttall (1818, v. 2, p. 58-59) in a new section named Gymnocaulis as Orobanche fasciculata, with calyx and corolla almost equally 5-cleft.
Original Text Comments 448. OROBANCHE. L. (Broomrape.) Calix 4 or 5-cleft, segments often unequal. Corolla ringent. Capsule ovate, acute, 1-celled, 2-valved; seeds numerous. A gland beneath the base of the germ. … [Description of the genus omitted.] … [Description of O. americana. omitted.] * Gymnocaulis. Calix and corolla almost equally 5-cleft. Assumed to be a new section name. 3. * fasciculata. Stem short and simple; peduncles many, naked, nearly terminal, and about the length of the stem; lobes of the corolla very short, rounded, and naked on the margin. Hab. With the above. Flowering in June and July. Very nearly allied to the following. The above is O. ludoviciana, which was “… in sandy alluvial soils, around Fort Mandan …” The following is O. biflora. Handwritten note, “primarily Artemisia.” Obs. 4 or 5 inches high, densely and pulverulently pubescent. Stem angular, with few scales, terminating in 6 or more naked filiform peduncles, 2 to 2 1-2 inches in length. Calix campanulate, semiquinquefid, segments sublanceolate, acute, little more than one-third the length of the corolla. Corolla curved, tubular, dilute purple; segments of the border scarcely more than a line long; lower lip producing a grooved palate. Stamina smooth, lobes of the anthers acute below, opening marginally and longitudinally, the lobes being parallel. Capsule 1-celled, ovate, acuminate.
Literature Cited:
- Gray, Asa, 1856.
Other articles:
• Taxa Notes:
Notes on Aphyllon, Gray, 1856;
Gray, Asa. 1856. 2nd ed. Manual of the botany of the northern United States. New York. Publication of Aphyllon fasciculatum
2. A. fasciculatum, Torr. & Gr. Scaly stem erect and rising 3'-4' out of ground, mostly longer than the crowded peduncles ; divisions of the calyx triangular, very much shorter than the corolla, which has rounded short lobes. (Orobanche fasciculata, Nutt.) — Islands in Lake Huron (Engelmann), and northward. May.
Other articles:
• Golden Checklist Flora:
Orobanche uniflora;
(Syn: Aphyllon uniflorum (L.) Torr. & A. Gray , Orobanche uniflora L. var. uniflora )
Literature Cited:
- Linne´, Carl von, 1753.
Linnaeus (1753, v. 2, p. 633) ...
Original Text 5. OROBANCHE caule unifloro. uniflora. Orobanche aut Helleborine affinis marilandica, caule nudo, unico in fummitate flore. Raj. fuppl. 595. Dentariae f. Anblato cordi affinis, flore pallide caeruleo. Gron. virg. 70. Gentiana minor aureo, flore fimplici amplo deflexo pallide flavefcente. Pluk. mant. 89. t. 348 f. 3. Aphyllon. Mitch. 25. Habitat in Virginia.
Other articles:
• Golden Checklist Flora:
Orthocarpus luteus;
Literature Cited:
- Nuttall, Thomas, 1818.
Other articles:
• Publication Details:
Nuttall, 1818, publication details;
Nuttall (1818, v. 2, p. 57) described a new genus Orthocarpus and added O. luteus to it.
Original Text Comments 446. *ORTHOCARPUS. † Calix tubular, semiquadrifid. Corolla bilabiate, closed; upper lip smaller, compressed, with the margin inflected; lower concave, obsoletely 3-toothed, unexpanded. Anthers unconnected; lobes unequal, divaricate. Capsule straight, elliptic-ovate, 2-celled, 2-valved, many-seeded, opening on both sides; dissepiment transverse. Seeds small, alated lunate margin. Annual; stem simple; leaves alternate, entire; flowers axillary, alternate, sessile, and bracteate; bractes divaricately trifid. Species. 1. O. luteus. Descript. Root tortuous, perpendicular, and fibrous. Stem simple, hirsutely pilose, terete. Leaves increasing in size upwards, alternate, sessile, lanceolate-linear, acute, entire, opaque, and as will as the bractes and calix shortly and somewhat viscidly pubescent, margins scabrous; bractes cuneate, divaricately trifid, about an inch long and the same in width, 3-nerved. Calix much shorted than the bractes, partly compressed, striated, subcampanulate, segments linear-lanceolate, acute. Corolla yellow, smooth; tube slender, about the length of the calix; both of the lips concave, inflected, and closed, the lower somewhat plaited and terminated by 3 minute dentures, so inconspicuous as the give the corolla the appearance of being destitute of an under lip. Stamina 4, small; filaments capillary, ingrafted upon the upper lip a little below the orifice, approximating by pairs unde the same lip; anthers pale, unusually small, 2-celled, distinctly and separately 2-lobed, lobes pubescent, not parallel, one acute-angularly diverging from below the summit of the other, almost exactly similar to the small lamdba (sic) of the Greek alphabet (λ). Style filiform, stigma simple, minute. Capsule elliptic ovate, obtuse, staight (sic) and pubescent, included within the calix, 2-celled, 2-valved, many-seeded, margins of the valves partly inflected; dissepiment transverse, or origining from the middle of the valves and seminiferous. Seeds small, more than 10, having an alated interrupted margin. Cotyledones 2, very small, upon the growing plant, oval. Hab. In humid situations on the plains of the Missouri, near Fort Mandan; very local. Flowering in July and August. Height about 12 or 14 inches. Growing in quantities. Flowers of a bright and uniform yellow, almost of the size and form of the common species of Melampyrum at first sight. In point of affinity it cannot be compared with any other genus, notwithstanding its marked distinction.
† The straightness of the fruit, readily distinguishing this genus from Melampyrum.Melampyrum is a genus of about 20 species of herbaceous flowering plants in the family Orobanchaceae known commonly as cow wheat. They are native to temperate regions of the Northern Hemisphere, though not found in Colorado. They are hemiparasites on other plants, obtaining water and nutrients from host plants, though they are able to survive on their own without parasitising other plants.
Other articles:
• Golden Checklist Flora:
Erythranthe floribunda;
(Syn: Mimulus floribundus Lindl. )
Literature Cited:
- Lindley, John, 1827.
Lindley (1827, v. 13, t. 1125) ...
A neat hardy annual, found by Mr. Douglas on moist rocks in the interior of the districts of the river Columbia.
Other articles:
• Golden Checklist Flora:
Erythranthe glabrata;
(Syn: Mimulus glabratus Kunth)
(Syn: Mimulus guttatus DC.)
Other articles:
• Golden Checklist Flora:
Collinsia;
Literature Cited:
- Nuttall, Thomas, 1817.
Nuttall (1817, v. 1, pt. 1, p. 189) ...
Description of Collinsia, a new genus of plants. By Thomas Nuttall. In the spring of 1810, during the course of an extensive journey into the north-western interior of the territories of the United States, I first became acquainted with the very singular and interesting plant which forms the subject of the present memoir. The specimens which I then obtained on the alluvial soils of the Alleghany and on the borders of lake Erie were finally lost. On arriving at St. Louis, near the confluence of the Missouri and Mississippi rivers, I found that Mr. John Bradbury, a botanist, had also detected this plant about the same time, on the banks of the Missouri and Mississippi, but I saw no specimen. In the spring of last year (1816), having undertaken a tour to the western states, I determined, if possible, again to collect this neglected plant, but after a journey of more than a hundred miles, for scarcely any other purpose, I arrived at Pittsburgh disappointed of my object. On descending the Ohio, however, nearly to Galiopolis, I at last recognised it on the more open alluvions of the river, withered and nearly past affording seed, accompanied by the Hesperis pinnatifida and the interesting Phalangium esculentum: from these seeds I have been at last fortunate enough to obtain the plant from which the accompanying drawing was taken by my friend, Mr. C. A. Le Sueur. This plant has for several years been known to Dr. Drake of Cincinati, in whose collection I saw specimens of it, and also with Dr. Short of Lexington, Kentucky, two gentlemen, who, amidst the avocations of an irksome profession, have still found leisure to cultivate some of the branches of natural science. I have dedicated this genus to the name of Zaccheus Collins, Esq. of Philadelphia, a gentleman, whose talents as a botanist and a mineralogist are deservedly acknowledged.
The article goes on to a formal description of Collinsia verna and includes a black and white drawing.
Other articles:
• Golden Checklist Flora:
Collinsia parviflora;
Literature Cited:
- Lindley, John, 1827.
“Received by the Horticultural Society from Mr. David Douglas, in 1827, by whome it was found in the vicinity of the river Columbia (Lindley, 1827).”
Other articles:
• Golden Checklist Flora:
Gratiola neglecta;
Literature Cited:
- Torrey, John, 1819.
Torrey (1819, p. 10 & 89) ...
Original Text 9. Gratiola. L. Hedge Hyssop. 1 aurea. Muhl. (Golden-Pert.) In sandy woods in moist places. New-Jersey. Long-Island, &c. ♃. Aug. yellow. 2 Virginica. L.
officinalis. Mx.
B. Caroliniensis. Pers.In inundated and moist places (rare.) ♃. Aug. 3. neglecta. Sp. nov. In similar situations. (common.) July-Aug. ♃. white and yellow. … Page 10. Gratiola neglecta, nobis : G. caule pubescente assurgente tereti ; foliis glabris lanceolatis pauci-dentato-serratis basi attenuatis connatisque : calycis foliolis aequalibus ; filamenta sterilia nulla. Root fibrous, perenial. Stem about six inches, branched at the base, round ; leaves broad-lanceolate, narrowed at the base and a little connate, smooth, rather obtuse. Peduncles varying in length, some of them shorter, and some longer than the leaves, the upper ones opposite. Calyx with obtuse lanceolate segments a little more than one third the length of the corolla ; bracts longer than the calyx, beset with glanduliferous hairs. Corolla with divisions rounded, the upper segment longer, and all of them emarginate ; limb white ; tube yellow, bent obliquely, smooth ; throat hairy. Filaments shorter than the tube. Stigma oblique, compressed. Capsule ovate, acutish, and about the length of the calyx.
This species approaches very near to G. Virginica, and may be the plant which Pursh has described as such. From the G. Virginica of Elliott it differs in several circumstances. Particularly in having a pubescent stem, and in the absence of abortive stamina. It has great affinity to Gratiola sphaerocarpa. Ell.
Literature Cited:
- Pennell, Francis W., 1921.
Other articles:
• Golden Checklist Flora:
Linaria canadensis texana;
Pennell (1921, p. 502) ...
Original Text 1a. Linaria canadensis texana (Scheele) Pennell, comb. nov. Linaria texana Scheele, in Linnaea 21: 4761. 1848. “Zwischen Houston and Austin [Texas] haufig:” Romer." Description sufficiently distinctive. Here distinguished primarily by roughness of seeds, which however varies from a few roughenings on the side to densely tuberculate. The wealth of Texan material at my disposal shows that each character of the contrasts previously outlined (in Contrib. U. S. Nat. Herb. 20: 323. 1920, and Proc. Acad. Nat. Sci. Phila. 71 : 258. 1920) between this and Linaria canadensis breaks down in numerous instances through the area where the two overlap. Consequently it seems more rational to consider texana a variety, its range and relation to the species roughly paralleling that of Veronica peregrina xalapensis (H. B. K.) Pennell. For an excellent illustration of Linaria candensis texana see the Botanical Magazine 63: pi. 3473. 1836. Open sand or sandy loam, frequently or usually a weed, common through the Coastal Plain, chiefly westward; occasional and probably introduced inland. Ranges widely through western North America, and into South America. List of specimens examined from Arkansas, Oklahoma, Louisiana, and Texas omitted.
Other articles:
• Golden Checklist Flora:
Linaria dalmatica;
(Syn: Linaria dalmatica (L.) Mill. ssp. dalmatica, Linaria genistifolia (L.) Mill. ssp. dalmatica (L.) Maire & Petitm.)
Literature Cited:
- Linne´, Carl von, 1753.
Linnaeus (1753, v. 2, p. 616) ...
Original Text dalmaticum. 18. ANTIRRHINUM foliis lanceolatis alternis, caule fruticofo. Linaria latifolia dalmatica, magno flore. Bauh. pin. 212. prodr. 106. Linaria maxima, folio lauri. Bauh. hift. 3. p. 458. Habitat in Creta. ♄.
Other articles:
• Golden Checklist Flora:
Penstemon secundiflorus;
Found in every Golden s.l. open space. Jefferson County collections along the base of the Front Range and barely into the foothills, with just a few in the interior of southern county. Most collections along the Front Range and interior valleys.
Name applied by Bentham (1846) to a specimen seen in Torrey's herbarium, possibly a Fremont collection.
Literature Cited:
- Bentham, George, 1846.
Bentham in DeCandolle (1846, v. 10, p.325) ...
25. P. secundiflorus, erectus, foliis radicalibus petiolatis ellipticis lanceolatisve acutis, superioribus anguste lanceolatis, panicula angusta, pedunculis erectis plurifloris secundis, calycis segmentis ovali-oblongis acutis margine membranaceis, corollae tubo superne ampliato, antheris glabris, filamento sterili apice dilato barbato. ♃ in montibus scopulosis (Fremont !). Hinc P. nitido, illinc P. coeruleo affines. Caules 9-10-pollicares. Cymae laxiusculae, superiores 1-3-florae, inferiores sub-5-florae. (v. in herb. Torrey).
Other articles:
• Golden Checklist Flora:
Penstemon strictus;
Literature Cited:
- Bentham, George, 1846.
Bentham in DeCandolle (1846, v. 10, p. 324) ...
Original Text Translation and Interpretation 24. P. strictus, rigidus, subglaucus, foliis imis oblongo-spathulatis petiolatis, caulinis lineari-lancelatis, panicula elongata angusta virgata , pedunculis brevibus plurifloris , calycis segmentis ovali-oblongis obtusis margine membranaceis, antheris parce piloso-lanatis , filamento sterili superne dilatato subnudo. ♃ in montibus Scopulosis ad fontes fl. Sweetwater (Fremont !). P. micranthus Torr. in Frem. rep. p. 89, non Nutt. Specimen vidi unicum sesquipedale. Folia quam in P. coeruloa angustiosa. Corollae imperfectae iis P. coerulei subsimiles videntur. Antherarum pili pauci, longi, molles (v. in herb. Torrey.) 24. P. tight, rigid, subglaucous, the lower leaves oblong-spathulate, petiolate, of a linear-lanceolate cauline, elongated panicle, narrow striped peduncles calyx with short, numerous, oval-oblong segments, obtuse with membranous margin, anthers sparsely hairy, woolly, with a sterile filament enlarged above, I subdo. [forb] In the Rocky Mountains at the Sweetwater River Springs (Fremont!). P. micranthus Torr. in Frem. rep. p. 89, non Nutt. I saw only one foot and a half specimen. Leaves as in P. coeruloa angustiosa. The imperfect corolla appear to be similar to those of P. coerulei Anther hairs few, long, soft (seen in Torrey herbarium).
Other articles:
• Golden Checklist Flora:
Penstemon virens;
The name P. virens was applied by Rydberg (1917) apparently at the suggestion of Pennell who collected the type. The type locality is the general area of Red Rocks Amphitheater though the amphitheater did not exist at the time. As an entity it was known well before then, as early as Nuttall's 1834 trip to Oregon Territory, and by Parry's collection, and Hall & Harbour collections. However, it was treated as a form of P. pubescens until treated as P. humilis Gray (1862), of which part was segregated into P. virens Rydberg (1917) based upon Pennell's type.
Literature Cited:
- Gray, Asa, 1862.
Gray (1862, p. 69-70) ...
43. P. humilis, Nutt. in herb. Acad. Philad. P. gracili Nutt. (P. pubescenti var. gracili supra) maxime affinis, sed vulgo humilior (3-9-pollicaris), foliis pallidioribus, thyrso etiam strictiore, corolla saturate caerulea breviore (haud ultra-semipollicari) fauce satis ampliata hiante plicis sub labio inferiore obsoletis. — Rocky Mountains, Nuttall (a very depauperate doubtless alpine specimen in herb. Acad. Philad.) ; common about Pike's Peak and vicinity, Parry, no. 257 (Enum. p. 27), Mr. Howard, and in taller specimens than before seen, Hall and Harbour, 1862, no. 387, on low mountains, “an early and very pretty species,” confirming its close relationship with P. gracilis. Yet this cannot be regarded as a variety of P. pubescens.
Literature Cited:
- Rydberg, Per Axel, 1917.
Rydberg (1917, p. 773 and 1066) ...
60. P. virens Pennell. Stem 1-3 dm. high, glabrous up to the inflorescence; leaves glabrous, glaucescent, the lower petioled; blades 2—4 cm. long; upper stem leaves lanceolate or oblong, usually denticulate; calyx 4 mm. long; corolla deep blue, 15 mm. long, funnelform; lower lip slightly bearded; sterile stamen yellow bearded on the spatulate tip. P. humilis A. Gray, mainly, not Nutt. Plains and hills: Alta. — Colo. — Nev. Plain — Mont. Je-Au. ... New Genera and Species Pentstemon virens Pennell. Type: Stony hillsides, foothills north of Morrison, Colo., 1915, Pennell 5821 (N.Y.). P. humile A. Gray, Am. Jour. Sci. II, 34: 253. 1862. Not Nutt.
Other articles:
• Golden Checklist Flora:
Penstemon virgatus var. asa-grayi;
Literature Cited:
- Crosswhite, Frank S., 1965.
Locations: Empire.
Crosswhite (1965, p. 318) ...
... I have recently concluded that the plant called “P. unilateralis” by authors is nothing more than a well-marked subspecies of P. virgatus A. Gray (in N+bot. Mex. Bound. surv. 113. 1859; isotype seen). Dr. William T. Penland has provided the only description of the subspecies in H. D. Harrington's Manual of the Plants of Colorado (Sage Books, Denver, 1954). In recognition of the excellence of Gray's revisions of the genus, this previously nameless Penstemon is here designated Penstemon virgatus ssp. Asa-Grayii Crosswhite, ssp. nov. — foliis radicalis oblanceolatis; a ssp. virgatus caulibus glabrus differt: typus: H. N. Patterson 258 (PH, isotypes NY, US, WIS) Empire, Clear Creek Co., Colo., Jul. 13, 1892. Mounted with this flowering material, in all 4 herbaria cited, is fruiting material collected in Sept. of the same year. &Mdash; Frank S. Crosswhite, Herbarium, Universoty of Wisconsin, Madison, 53706.
Other articles:
• Golden Checklist Flora:
Plantago lanceolata;
Other articles:
• Golden Checklist Flora:
Plantago major;
Literature Cited:
- Linne´, Carl von, 1753.
Linnaeus (1753, v. 1., p. 112-113) ...
Original Text major. 1. PLANTAGO foliis ovatis glabris, nudo fcapo tereti, fpica flofculis imbricatis. Plantago foliis ovatis glabris. Hort. cliff. 43. Fl. fuec. 122. Mat. med. 49. Roy. lugdb. 401. Hall. helv. 470. Dalib. parif. 50. Plantago fcapo fpicato, foliis ovatis. Fl. lapp. 62. Plantago latifolia finuata. Bauh. pin. 189. Plantago major. Cam. epit. 261. β. Plantago latifolia glabra minor. Bauh. pin. 189. γ. Plantago latifolia rofea, floribus quafi in fpica difpofitis. Bauh. pin. 189. Hort. upf. 29. δ. Plantago latifolia rofea, flore expanfo. Bauh. pin. 189. ε. Plantago latifolia, fpica multiplici fparfa. Bauh. pin. 189. Habitat in Europa ad vias. ♃.
Other articles:
• Golden Checklist Flora:
Plantago patagonica;
Jacquin (1793, v. 2., p. 9) short description with a reference to Jacq. coll. vol. 5. [Collectaneorum supplementum, where there is a longer description and a collecting locality.] and Jacquin (1793, v. 2, t.(plate) 306.
Literature Cited:
- Jacquin, Nikolaus Joseph, 1796.
Jacquin (1796, p. 35) ...
Original Text Translation and Comments Crefcit ad Champion-River in Patagonia. Grows at Champion-River in Patagonia.
Other articles:
• Notes on Colorado Flora:
Synthyris plantaginea;
Literature Cited:
- James, Edwin, 1825.
James, Edwin. 1826. Catalogue of Plants collected during a Journey ... Transactions of the American Philosophical Society. new ser.:v.2 (1825)
V[eronica] plantaginea. N. S.? Flowers spiked, unibracteate. Capsule nearly orbicular, dehiscent at the margin. Radical leaves numerous and large, oval-lanceolate, stiped. Margin crenate. Those of the scape few, sessile and alternate. Spike long, dense-flowered. This plant, so unlike in habit and appearance to the American Species of Veronica, seems referable to no other Genus among the Pediculariae of Jussieu, to whic it is manifestly nearly allied. Habitat on open plains within the Rocky Mountains; flowering in June and July.
Literature Cited:
- Rydberg, Per Axel, 1903.
Rydberg, Per Axel. 1903. Some Generic Segregations.
IV. A fourth segregation is proposed in Synthyris. Professor Greene has included this genus in Wulfenia and he has been followed by many American botanists. Wulfenia, represented by European and Asiatic plants, has a 4-valved capsule, which is neither flattened nor emarginate at the apex, and a corolla with along tube. All the American species have flattened capsules, which are 2-valved and obcordate at the apex. In this respect they agree perfectly with Veronica. The original Synthyris (S. reniformis) and a few related species have a corolla that is built exactly on the same plan as that of Veronica, and it is only the general habit that keeps them out of that genus. S. alpina and its relatives, on the contrary, have an altogether different corolla, cleft to near the base into two distinct lips. S. rubra and its allies have no corollas but agree otherwise both in general appearance and structure of the flower and fruit with S. alpina. It would however hardly be advisable to place them in different genera merely on account of the presence or absence of the corolla. The related genera are distinguished as follows : 1a. Corolla scarcely 2-lipped ; almost equally deeply 4-lobed. 2a. Corolla tubular-funnelform; capsule neither flattened nor obcordate at the apex, 4-valved. 3a. Leaves alternate but mostly basal. Wulfenia. 3b. Leaves opposite; stem leafy. Leptandra. 2b. Corolla rotate or short-campanulate ; capsule flattened, obcordate, 2-valved. 4a. Leaves opposite or whorled ; stem leafy. Veronica. 4b. Leaves alternate, but mostly basal ; plant with a mostly naked scape, Synthyris. 1b. Corolla, if present, deeply 2-lipped; upper lip entire, broad, arcuate ; lower lip straight and deeply laciniate ; leaves alternate but mostly basal ; scape bracted. Besseya.Synthyris Benth.; DC. Prodr. 10: 454. 1846 This genus was based on S. reniformis. S. rubra was also included in the genus by the author, but he admits that it is a rather anomalous member, lacking the corolla. It is, however, more nearly related to S. alpina, which has a corolla of a different structure. The species belonging here are the following : S. renitformis (Dougl.) Benth., S. major (Hook.) Heller, S. pinnatifida S. Wats., and S. laciniata (Gray) Rydb. Whether S. rotundifolia A. Gray should also be included or should be made the type of a separate genus is a little doubtful. At present I am inclined to regard it as a Synthyris. So also S. schizantha Piper, with lacerate corolla-lobes, which species is still more aberrant. Besseya gen. nov. Low perennials with a thick rootstock. Basal leaves petioled ; blades ovate or oblong, crenate; stem-leaves bract-like, alternate: inflorescence a bracted spike: calyx almost regularly 4-cleft to near the base or cleft to the base on the upper side and then 2—3- lobed : corolla 2-lipped, cleft to near the base, or lacking; upper lip, if present, entire, obovate or cuneate-obovate, concave ; lower lip much shorter, often irregularly 2-3-cleft or laciniate : stamens 2, attached at the base of the corolla, or if this is absent on a small disk: capsule flattened, obcordate, 2-valved. It is with pleasure that I dedicate this genus of western mountain plants to Professor Charles E. Bessey, of the University of Nebraska, my former teacher and the most prominent botanist of the region where these small plants grow. The first species cited below is regarded as the type.
a. With corolla:
1. B. alpina (A. Gray); Syuthyris alpina A. Gray, Am. Jour. Sci. II 34; 251. 1862.
2. B. Bullii (Eaton) ; Gymnandra Bullii Eat.; Eat. & Wr. N. Am. Bot. 259. 1840; Synthyris Houghtoniana Benth.; DC. Prodr. 10: 454. 1846.
3. B. plantaginea (Benth.); S. plantaginea Benth.; DC. Prodr. 10: 455. 1846.
4. B. reflexa (Eastw.); S. reflexa Eastw. Proc. Cal. Acad. III. 1: 124. 1398.
5. B. Ritteriana (Eastw.) ; S. Ritteriana Eastw. l. c. 123.
b. Without corolla :
6. B. rubra (Dougl.); Gymnandra rubra Dougl.; Hook. Fl. Bor. Am. 2: 103. 1838; Synthyris rubra Benth.; DC. Prodr. 10: 455. 1846.
7. B. gymnocarpa (A. Nels.); Wulfenia gymnocarpa A. Nelson, Bull. Torrey Club, 25: 282. 1898.
8. B. Wyomingensis (A. Nels.); W. Wyomingensis A. Nelson, l. c.281.,
Literature Cited:
- Albach, Dirk C., 2019.
28 Bract margins: proximals serrate, distals sometimes entire; leaf blades (6–)12–28(–35) mm; styles (0.7–)1–1.3(–1.7) mm. Veronica argute-serrata 28 Bract margins entire; leaf blades (3–)4–12(–20) mm; styles 0.4–0.8(–2) mm. Veronica biloba
Other articles:
• Golden Checklist Flora:
Veronica americana;
Literature Cited:
- Bentham, George, 1846.
Bentham in DeCandolle (1846, v. 10, p. 468) ...
Original Text 47. V. Americana (Schweinitz! mss.), glabra, basi decumbens , superne erecta, foliis plerisque petiolatis ovatis oblongisve acutis vel obtusiusculis serratis crassiusculis basi truncato-subcordatis, capsula subrotunda turgida emarginata. ♃ in America boreali a Canada et Carolina usque ad flum. Oregon et in ins. Sitcha. V. Anagallis auct. plur. amer. non Linn. Species inter V. Anagallidem et V. Beccabungam fere media. Habitus potius prioris sed folia pleraque petiolata, breviora, latiora. Capsula et semina V. Beccabungae. Folia nunc vix semi-pollicaria, nunc 1-2poll. longa. (v. s. )
Bentham wrote the section on Scrophulariaceae.
Other articles:
• Golden Checklist Flora:
Veronica anagallis-aquatica;
Literature Cited:
- Linne´, Carl von, 1753.
- Stearn, William T., 2004.
Linnaeus (1753, v. 1, p. 12) ...
Anagall. ᐁ 15. VERONICA racemis lateralibus, foliis lanceolatis ferratis, caule erecto. Fl. fuec. 10. Dalib. parif. 7.   Veronica foliis oppofitis laevibus crenatis, floribus laxe fpicatis ex alis. Gron. virg. 4. Veronica aquatica major, folio oblongo. Bauh. pin. 252. Habitat in Europa ad foffas. ☉
The upside-down triangle (ᐁ) is a symbol for water, or aquatica, e.g., Anagall. ᐁ = Anagallis-aquatica (Stearn, 2004). Linneaus (1753) used symbols forming part of a specific epithet under Alisma, Campanula, Cotyledon, Scandix, Sysimbrium, and Veronica.
Literature Cited:
- Ackerfield, Jennifer, 2015.
Ackerfield (2015) treats V. anagallis-aquatica as a native, and places V. catenata in synonomy.
Literature Cited:
- POWO, 2021 - 2026.
Kew POWO maps V. anagallis-aquatica as introduced to Colorado.
Literature Cited:
- Albach, Dirk C., 2019.
Veronica argute-serrata has been recorded mostly under the name V. biloba and occasionally under V. campylopoda Boissier. Most of the records for V. biloba likely belong under this name. Veronica biloba is also found in the flora area, sometimes mixed with V. argute-serrata; V. campylopoda has not been recorded from the flora area. A chromosome count of 2n = 18 (C. R. Bell 1965) for American material is suspicious because 2n = 42 has been reported multiple times in Turkish plants, and a base chromosome number of × = 9 is not known in related species.
Other articles:
• Golden Checklist Flora:
Veronica arvensis;
Literature Cited:
- Linne´, Carl von, 1753.
Linnaeus (1753, v. 1, p. 13) ...
Original Text 21. VERONICA floribus folitariis, foliis cordatis incifis pedunculo longioribus. Fl. fuec. 16. Dalib. parif. 5. arvenfis Veronica foliis oppofitis cordatis crenatis, floribus folitariis feffilibus. Hort. cliff. 9. Gron. virg. 4. Roy. lugdb. 303. Alfine veronicae foliis, flofculis cauliculis adhaerentibus. Bauh. pin. 250. Habitat in Europae arvis, cultis. ☉.
Other articles:
• Golden Checklist Flora:
Veronica biloba;
(Syn: Pocilla biloba (L.) W. A. Weber)
Literature Cited:
- Linne´, Carl von, 1771.
Linnaeus (1771, p. 172) ...
Original Text Translation and Interpretation Habitat inter Cappadociae fegetes. D. Schreber. ☉. She lives among the crops of Cappadocia [central Turkey]. D. Schreber. Annual.
Other articles:
• Golden Checklist Flora:
Veronica catenata;
Literature Cited:
- Pennell, Francis W., 1921b.
Pennell (1921, v. 23, n. 266, p. 37) ...
Original Text 27. Veronica catenata Pennell, sp. nov. Flowering stem 1-3 dm. long, glabrous throughout. Leaves lanceolate, acute or acutish, crenate to nearly entire, 3-5 cm. long, 1 cm. wide, all clasping, when submersed elongating and reaching 12 cm. long and 2 cm. wide. Racemes axillary to the upper leaves, 6-12 cm. long, 15-25-flowered. Bracts narrowly lanceolate, 4-5 mm. long. Pedicels 3-5 mm. long, glabrous. Sepals 3-3.5 mm. long, lance-ovate, obtusish. Corolla-lobes pale-blue. Style 1.2-1.7 mm. long. Capsule 3 mm. long, 3.5 mm. wide, broad-globose, decidedly emarginate. Seeds 0.5 mm. long, yellow-brown. Type, Hot Springs, South Dakota, collected in flower and fruit June 16, 1892, P. A. Rydberg 926, in Herb. New York Botanical Garden. Named from the chain-like aspect of the long racemes of short-pedicelled flowers. Slow-flowing streams, plains, from North Dakota and Saskatchewan to Kansas and New Mexico, southward west to Nevada and southern California.
Literature Cited:
- Weber, William A., and Ronald C. Wittmann, 2012.
Accepted by Weber & Wittmann (2012) in a paragraph that also mentions V. anagallis-aquatica noting that it is closely related.
V. catenata is not identified as alien.
Literature Cited:
- Ackerfield, Jennifer, 2015.
Treated by Ackerfield as a synonym of V. anagallis-aquatica.
Literature Cited:
- POWO, 2021 - 2026.
Kew Plants of the World Online accepts both V. anagallis-aquatica and V. catenata.
According to Kew POWO, V. catenata does not occur in Colorado, and V. anagallis-aquatica is introduced to Colorado.
Other articles:
• Golden Checklist Flora:
Veronica peregrina ssp. xalapensis;
(Syn: Veronica xalapensis Kunth, V. peregrina L. var. xalapensis> (Kunth) H.St.John )
Literature Cited:
Kunth, Nov. Gen. Sp. [H.B.K.] 2(pt. 8): 389 (ed. qto.) (1818).
Literature Cited:
Pennell (1919, v. 19, n. 9, p. 167) ...
Other articles:
Galium aparine L., Sp. Pl. 1: 108 (1753).
Other articles:
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Griesb., 1839. Spicilegium florae rumelicae et bithynicae exhibens synopsin plantarum. 2. p. 178
Other articles:
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Literature Cited:
Gray (1856, p. 305) in his preface to the list of plants collected in Japan ...
Gray (1856, p. 313) describes our plant ...
Other articles:
Literature Cited:
Linnaeus (1753, v. 1, p. 173-174) ...
In Linnaeus' time, “Tataria” referred to Central Asia and European Russia east of the river Don (Stearn, 1992).
Literature Cited:
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Hooker (1840, v. 1, p. 285), volume 1 being concluded in 1834. IPNI shows the date of publication to be 1833.
Other articles:
Literature Cited:
Gray (1853, v. 5, p. 66) writing in Plantae Wrightianae described S. rotundifolius
Other articles:
Literature Cited:
Linnaeus (1753, v. 1, p. 268) ...
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Locations:
Schweich Hill.
There is one collection of “Dane's Blood” — Campanula glomerata L. —
in Golden s.l. that was made on Schweich Hill.
It is not really a garden escapee, since it was clearly thrown or dumped in the open space, i.e., it did not escape a garden on its own.
Perhaps the correct term is that it was “yeeted” into the open space.
It is the only collection from Jefferson County.
There are three other collections scattered around Colorado, all of which appear to be garden escapees.
It is one of four unrelated taxa known commonly as “Dane's Blood,” having in folklore sprung up in fields where Danes were slaughtered.
The plant was described by Linnaeus (1753) who noted that it lived in England, France, and Sweden.
Literature Cited:
Linnaeus (1753, v. 1, 166-167) ...
Literature Cited:
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Literature Cited:
Linnaeus (1753, v. 1, 165) ...
Other articles:
Linnaeus (1753, v. 1, p. 163) ...
Literature Cited:
In a paper designed to explore the roles that glaciation and whole genome duplication may have played in the evolution of C rotundifolia, Sutherland & Galloway (2018) constructed a chloroplast phylogeny of 37 populations that spanned the range of the C. rotundifolia complex.
While they showed two European clades and one North American clade were likely genetically isolated from each other, and that the North Amrican clade was likely sourced from a glacial refugium in midwestern North America.
To date, this is the only paper I have found showing how the North American members of the C. rotundifolia complex are sufficiently phylogenetically distinct to justify taxonomically isolating them at the species rank, i.e., as C. petiolata.
Literature Cited:
“Thistles” in the broad sense are in the
tribe Cardueae Cassini, J. Phys. Chim. Hist. Nat. Arts. 88: 155. 1819 .
Tribe Cynareae Lam. & DC. would seem to have precedence over Cardueae by nearly a decade.
The North American representatives of Tribue Cardueae accepted by FNANM are as follows.
Taxa found in Colorado are shown in bold.
There is a more recent paper (Herrando-Moraira, Sonia, et al. 2019) describing the phylogeny of Cardueae using Hyb-Seq data.
It is, however, behind a paywall until August 1, 2020, and we eagerly await its unveiling.
Other articles:
Literature Cited:
Linnaeus (1753, v.2, p. 899) published Achillea millefolium without mention of
habitat in North America.
Habitat in Europć pascuis pratisque.
♃
Literature Cited:
Kalm (1762, v. III., p. 499) notes presence in Montreal, Canada,
of A. millefolium, in addition to Prunella vulgaris, Oenothera biennis,
Viola canadensis, and others.
There might be additional information but I struggle to read Swedish in the old font
used in the report.
Literature Cited:
Pursh (1814, v. 2, p. 563) thought that Achillea millifolium
may have been introduced from Europe.
Icon. Fl. dan. 737.
Engl. bot. 758.
In fields an on road sides ; common.
♃.
June-Aug.
v. v.
Probably introduced from Europe.
Flowers white or rose-coloured.
Literature Cited:
Nuttall (1818, v. 2, p. 171) also thought that A. millefolium was naturalized in North America.
Literature Cited:
Torrey & Gray (1841-1843, v. 2, p. 409) recognized that A. millefolium occurred throughout North America, but also noted it was introduced from Europe into pastures, &c.
[Greek agos, leader, and seris, chicory; allusion unclear]
Agoseris has a New World, amphitropical distribution.
All of the species are restricted to North America except A. coronopifolia (D’Urville) K. L. Chambers,
which is found in temperate regions of southern South America.
The South American disjunction appears to be the result of long-distance dispersal from North America (K. L. Chambers 1963).
Literature Cited:
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Schreber (1789-1791) published edition 8a of Linnaeus' Species Plantarum.
In the second volume he described Krigia Schred.
The volume is not available online, so I am unable to read the description.
Publication of this volume preceeded the publication of Troximon by Gaertner.
How much it preceeded Gaertner's publication, I don't know.
But it was enough to give Krigia priority over Troximon.
Literature Cited:
C. C. Robin published an account of his travels in Louisiana, West Florida and the West Indies from 1802 to 1806.
He included descriptions of plants observed in Louisiana.
Rafinesque undertook to translate, revise, and improve (as he says!) the plant list, and to see it published in New York.
Literature Cited:
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Literature Cited:
Phylogenetic analysis of relationships in the Cichorieae place Agoseris in the Microseris clade along with Nothocalais, Uropappus, and Stebbinsoseris.
Krigia (and Glyptopleura) was not placed in any major clade.
Other articles:
Troximon glaucum Pursh, Fl. Amer. Sept. (Pursh) 2: 505 (1814).
Literature Cited:
Other articles:
Pale Goat-Chicory was first published by Frederick Pursh in Volume 2 of his Flora Americae Septentrionalis.
What we now consider to be a male symbol – ♂ – probably means the plant is biennial.
Agoseris glauca (Pursh) Raf., Atlantic J. 6: 39 (1833).
Other articles:
T. parviflora [=A. parviflora] will be segregated into the second section that has a distinct rostrum (beak).
Literature Cited:
Other articles:
Though Rafinisque published Agoseris in 1817,
it wasn't until 1847 that D. Dietrich placed Pursh's Troximon glaucum in Agoseris.
Other articles:
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Nuttall (1841) described a new species of Troximon, T. parviflorum in describing the plants from his travels along the
Oregon Trail in 1834 and 1835.
It will become our current Agoseris parviflora.
Literature Cited:
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Dietrich (1847) in his Synopsis Plantarum placed the North American Troximon into Agoseris Raf.
Other articles:
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Other articles:
Literature Cited:
DeCandolle (1836, v. 5, p. 526) published A. psilostachya
from a collection by Berlander in Mexico.
Other articles:
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Linnaeus (1753, v. 2, p. 850) was fairly clear that G. margaritaceum
was native to North America.
Gnaphalium latifolium americanum. Bauh. pin. 263.
Gnaphalium americanum. Clif. hift. 1. p. 327.
Habitat in America feptentrionali, kamtfchatca.
♃
Literature Cited:
DeCandolle (1837, v. 6, p. 271) published Anaphalis, but he placed our margaritacea in Antennaria Sect. II. Margaripes.
Literature Cited:
Weber & Wittman (2012) state that Anaphalis is an anagram of Gnaphalium.
Other articles:
Literature Cited:
Other articles:
Nuttall (1841) first described A. parvifolia collected on his trip on the Oregon Trail with the Wyeth expedition.
Literature Cited:
Randal J. Bayer, who wrote the treatment of Antennaria for the Flora of North America
published a cladistic analysis (Bayer, 1990) of the sexually reproducing Antennaria.
However, A. parvifolia is not one of them because some individuals are dioecious and others are apomictic.
The article is behind a paywall.
I thought this was interesting, “ … polyploidy and two asexual means of reproduction, agamospermy and horizontal stolons,
have evolved in the more specialized groups, the Pulcherrimae and Catipes … ”
This would imply to me that A. parvifolia would be a member of one of those two groups.
Other articles:
Literature Cited:
Greene (1898, p. 281) in his “Sect. 2. Some northern species of Antennaria”
described a new Antennaria.
Other articles:
Literature Cited:
Hooker (1834, v. 1, p. 334) described A. cordifolia from
collections by Drummond in the Rocky Mountains and by Douglas along the Columbia River in Washington Territory.
Other articles:
Literature Cited:
Pursh (1814, v. 2, p. 527) published A. fulgens
from an unidentified collection on the banks of the Missouri.
There is no Lewis & Clark collection of this taxon,
so it may have been an unattributed Bradbury or Nuttall collection.
On the banks of the Missouri.
♃.
v. s. About a foot
or more high, very slender ; leaves on the stem generally
two pairs ; flowers somewhat smaller than the
preceding, of a very deep and beautiful yellow.
The preceding was A. plantaginea which Nuttall will reduce to a variety of A. fulgens.
Literature Cited:
Nuttall (1818, v. 2, p. 164) treated A. fulgens
as a variety of A. montana.
Literature Cited:
Riddell (1835, p. 52) ...
Artemisia L. is the largest genus of family Asteraceae, comprising about 350–500 diverse species. Artemisia has two basic chromosome numbers, with ploidy levels x=9 and x=8. Chromosome number in diploid is most often 2n=18 or 16. The genus is divided into five large groups Absinthium DC., Artemisia L., Dracunculus Besser, Seriphidium Besser and Tridantatae (Rydb.) McArthur. Its phylogeny was based on the two hypothesized evolutionary trends, loss of fertility in the disc florets and loss of ray florets.
Four taxa of Artemisia within subgroup Absinthium are recognized by Ackerfield (2015).
Literature Cited:
Eight taxa of Artemisia within subgroup Artemisia are recognized as occurring in Colorado (Ackerfield, 2022).
Rhizomatous
Literature Cited:
Four species of Artemisia within subgroup dracunculus are recognized by Ackerfield (2015).
Five secies are not recognized by Ackerfield (2015):
Literature Cited:
Eight taxa of Artemisia subg. Tridentatae are recognized as occurring in Colorado (Ackerfield, 2022).
Other articles:
Literature Cited:
Other articles:
Literature Cited:
Linnaeus (1753, v. 2, p. 846) published A. campestris stating that its habitat was dry, sunny fields in Europe.
Abrotanum campeftre. Bauh. pin. 136.
Ambrofia altera. Cam. epit. 597.
Habitat in Europć campis apricis, aridis.
♃
Habitat in sunny, dry European fields.
Literature Cited:
Michaux (1803) Flora boreali-americana, vol. 2 … published A. caudata at the rank of species.
Literature Cited:
Nuttall (1841, p. 401) in
“Descriptions of new species and genera of plants in the natural order of the Compositae, ...”
... published Artemisia pacifica at the rank of species.
Literature Cited:
Hall & Clements. 1923.
The Phylogenetic Method in Taxonomy: The North American Species of Artemisia, Chrysothamnus, and Atriplex is a 1923 publication by Harvey M. Hall and Frederic E. Clements, published by the Carnegie Institution of Washington, that applies phylogenetic principles to classify these three North American plant genera, focusing on evolutionary relationships rather than just morphology. The book is a significant work in botanical taxonomy, available in modern reprints and digital archives, and serves as a reference for understanding evolutionary classification.
Hall & Clements (1923) published … Artemisia campestris ssp. pacifica, i.e., at the rank of subspecies …
(A. pacifica Nuttall, Trans. Am. Phil. Soc. II, 7:401, 1841.)
Type locality, shores of the Pacific at the outlet of the Oregon (Columbia River), in sandy places.
The common form in western United States:
South Dakota and western Nebraska to New Mexico, Arizona, Oregon, Yukon, and Saskatchewan.
Collections:
Brookings, South Dakota, September, 1894, Thornber (UC);
Laramie, Wyoming, Nelson 7983 (Gr, NY, UC, as A. camporum Rydberg, minor variation 7);
Kiowa Valley, western Nebraska, Rydberg 203 (NY, same variation) ;
Ruxton Dell, Pikes Peak, Colorado, Clements 156 (DS, Gr, NY, US, same variation);
La Sal Mountains, Utah, Purpus 7019 (UC);
near Silver City, New Mexico, October 8, 1880, Greene (UC);
San Francisco Mountains, Arizona, Leiberg 5840 (US);
Bingen, Klickitat County, Washington, Suksdorf 4602 (DS, US);
Vancouver Island, British Columbia, August 16, 1887, Macoun (NY, US);
Clarks Fork Valley, northern Idaho, Leiberg 1580 (Gr, UC, NY, US);
Bozeman, Montana, Blankinship 304 (hb. Blankinship) ;
Whiteshore Lake, Saskatchewan, Macoun and Herriot 72832 (NY, minor variation 7, A. camporum Rydberg).
Hall & Clements (1923) published … A. campestris ssp. caudata at the rank of subspecies …
Neither I nor ChatGPT were able to find an earlier suggestion of this taxonomic change was published.
(A. caudata Michaux, Fl. Bor. Amer. 2:129 1803.)
Type locality, sandy banks of the Missouri River.
Widely,
distributed but most common in the eastern and central States: New Brunswick and
Maine to Florida, Texas, Wyoming, Washington, Saskatchewan, and Ontario.
(Specimens seen from south of Connecticut and Illinois are without roots and their identification
therefore not absolutely positive.)
Collections:
Pine Point, Maine, Parlin 1097 (N. E. Bot. Club);
Lynn Beach, Massachusetts, August, 1887, Summers (US);
North Haven, Connecticut, Bissell 878 (Gr);
Hennepin County, Minnesota, August, 1890, Sandberg (UC);
Washington Island, Wisconsin, September 12, 1889, Shuette (Gr, UC);
sandy barrens near Oquawka, Illinois, August, 1877, Patterson (UC);
sand dunes at Millers, Indiana, September 4, 1911, Sherff (US);
Blue River, Nebraska, September, 1888, Wibbe (UC);
vicinity of Pine Grove, Colorado, Crandall 2640 (US);
Deadwood, South Dakota, Carr 29 (NY, minor variation 18, A. forwoodi Watson);
Willow City, North Dakota, September 11, 1899, Lunell (NY, same variation);
near Bingen, Washington, August, 1907, Suksdorf (NY);
Brandon, Manitoba, Macoun 12247 (NY).
Literature Cited:
Palmer & Steyermark.
1935.
An Annotated Catalogue of the Flowering Plants of Missouri.
Annals of the Missouri Botanical Garden.
pp- 375-758,
p. 676
...
placed A. caudata as a variety of A. campestris rather than a subspecies.
Literature Cited:
Peck, Morton Eaton
A manual of the higher plants of Oregon.
p. 768 ...
placed A. pacifica as a variety of A. campestris rather than a subspecies.
Literature Cited:
The Flora of North America (Flora of North America Editorial Committee, eds., 2006) was written by Leila M. Shultz.
There is a local connection in that many collections at Rocky Flats were made by Ms. Shultz, who was working with G. Kunkel.
In her description of A. campestris, Ms. Shultz states that the species varies, and that each morphological form grades into another.
Taking a conservative approach, she recognizes three subspecies: subsp. pacifica (Nuttall) H. M. Hall & Clements, subsp. canadensis (Michaux) Scoggan, and subsp. caudata (Michaux) H. M. Hall & Clements.
Subspecies caudata and pacifica are known to occur in Colorado.
Literature Cited:
Other articles:
Weber & Wittmann (2012, p. 104) do not accept presence of A. campestris L.
in Colorado.
Instead, they accept Oligosporus caudatus (Michx.) Poljakov
and O. pacificus (Nutt.) Poljakov.
Literature Cited:
Ackerfield (2015, p. 121, and 2022, 2nd ed., p. 138) accepts three varieties of A. campestris L.
in Colorado, variety purshii (Hook.) Cronquist, variety caudata (Michx.) Palmer & Steyerm., and variety pacifica (Nutt.) M. Peck.
Other articles:
See also: Artemisia groenlandica Hornem.
Literature Cited:
Pursh. 1814. Vol. II, p. 522 ...
Literature Cited:
Hooker, William Jackson, Sir, (1785-1865)
1840.
Flora boreali-americana, or, the botany of the northern parts of British America : compiled principally from the plants collected by Dr. Richardson,
pt. 12, p. 241-end.
p. 326
...
Literature Cited:
(A. borealis Pallas, Reise 3:755, 1776.)
Type locality, Siberia.
Greenland
and Hudson Bay to Vermont, western Ontario, Colorado (?), and Washington, thence
probably far to the north and northwest, since it is reported from Kotzebue Sound
(Hooker, Fl. Bor. Am. 1:327, 1833) and is not rare is Siberia.
Collections:
Grand Falls, Newfoundland, on ledges and talus, Fernald and Wiegand 6388 (DS, Gr, UC);
Gaspé County, Quebec, on serpentine detritus, Collins and Fernald 145 (NY, UC, US);
Gaspé County, Quebec, on cliffs, Collins, Fernald, and Pease 6183 (UC);
Mount Mansfield, Vermont, Eggleston 2027a (NY);
Mount Harvard, Colorado, Clements 31 (NY, referred here because of the glabrous involucres,
but doubtless a local derivative of subspecies spithamaea);
near Banff, Alberta, McCalla 2016 (US, type of A. maccallae Rydberg, minor variation 22);
White Horse Rapids, Yukon River, Macoun 79015 (NY, same variation).
Literature Cited:
Cronquist. 1950. Notes on Compositae of Northwestern United States
made a new combination in Artemisia …
Literature Cited:
Schulz (2006, v. 19, p. 506) accepts Artemisia borealis Pallas subsp. borealis...
Literature Cited:
Weber & Wittmann (2012) treat our plant as Oligosporus groenlandicus (Hornem.) A.Löve & D.Löve.
They also treat Artemisia campestris ssp. caudata as Oligosporus caudatus (Michaux) Poljakov
and subspecies pacifica as O. pacificus (Nuttall) Poljakov.
Other articles:
Literature Cited:
Other articles:
Linnaeus (1753, v. 2, p. 849) ....
Abrotanum lini folio acriori & odorato. Tournef. inft. 459.
Dracunculus hortenfis. Bauh. pin. 98.
Draco herba. Dod. pempt. 709.
Habitat in Sibiria, Tataria.
♃
Literature Cited:
Other articles:
Weber & Wittman (2015, p. 103) recognize
Oligosporus dracunculus (L.) Poljakov subsp. glaucus (Pallas) Löve & Löve for
our Artemisia dracunculus.
Literature Cited:
Ackerfield (2015, p. 121) accepts A. dracunculus without infraspecific taxa.
Other articles:
Literature Cited:
In describing the collections of Edwin James, MD, John Torrey (1828, v. 2, p. 211) informally “published” A. filifolia.
Obs. Besides the above six species, Dr. James found another,
which appears to be new, but the specimen is without
lower leaves, and cannot, therefore, be certainly determined.
Literature Cited:
Goodman & Lawson (1995, p. 159) note that James could have collected A. filifolia in any one of five states the expedition passed through, and that the species occurs abundantly in the Texas panhandle which is perhaps the most likely collecting site.
Other articles:
Literature Cited:
Willdenow (1803v. 3, pt. 3, p. 1838) published A. fridiga ...
I think the location of “Dauuriae” is likely equivalent to
“Dauria,” “Dahuria,” or “Davuria,” which Stearn (2004) cites as a name for a region of
southeast Siberia.
Other articles:
Taxa I requested to SEINet:
Taxa included by SEINet:
Literature Cited:
Friis, Ib. 1996, The Icones ... florae Danicae ... (1761-1883) as a source of names of flowering plants.
Acta Univ. Ups. Symb. Bot. Ups. 31:3, 7-25. ISBN 91-554-3707-9. Available on ResearchGate.
The paper is apparently: Bocher et al. (1978). Gronlands Flora. 3. ed. Kobenhavn
Literature Cited:
Bjork, Curtis R. (2012) …
Literature Cited:
Other articles:
Thomas Nuttall collected the type material of Artemisia ludoviciana
on the banks of the Mississippi River and on the alluvial plains of the Missouri in 1811.
This material still exists in the herbarium of
Academy of Natural Sciences of Drexel University (ANSP:PH) and can be found online:
https://swbiodiversity.org/seinet/collections/individual/index.php?occid=30011291
Both collections appear to be designated as type material.
Nuttall (1818, p. 143) published his taxon in Genera of North American Plants.
The specific epithet “ludoviciana” is a Latinization of “Louisiana.”
A. gnaphalodes Nuttall. is now treated as a synonym of A. ludoviciana Nuttall.
A. cernua Nuttall. is now treated as a synonym of A. dracunculus L.
Literature Cited:
Hab. In the central chain of the Rocky Mountains, in Thornberg's Pass, near the great passage to the plains of the Oregon.
At first sight it somewhat resembles some variety of A. vulgaris, but is very distinct.
Remarkable for its smoothness.
Height one to two feet; segments of the leaves a line wide.
Thornberg's Pass was named for the member of the party who assured Wyeth he had crossed the mountains by this route.
The party was on or near Hyndman Peak (12,078 ft.) (Graustein, 1967, p. 303, & Note 34, Page 443.).
Townsend (1839, p. 120-121) wrote about this day …
Wyeth's party exited from the mountains onto the Camas Prairie on the Camas Branch of the South Fork of the Boise River.
Literature Cited:
Other articles:
Locations:
Timberline Station.
(Clausen, Keck, and Heisey, 1940) published variety incompta (Nuttall) Keck in their study of the Artemisia vulgaris complex.
Clausen, Keck, and Heisey (1940) continue with this discussion of ssp. ludoviciana
Literature Cited:
Other articles:
Literature Cited:
Locations:
Lewis River.
Nuttall (1841, p. 398) proposed Artemisia tridentata ...
Hab. Plains of the Oregon, and Lewis’ River.
A low, but rather stout shrub, white with a close tomentum.
Leaves rather more than an inch long, about two lines wide, more or less deeply
three-toothed, sometimes entire, the upper ones always so.
Panicle much branched, the flowers small.
(I have not seen them in a perfect state, and therefore class this species by its apparent
affinity with the last.)
Somewhat allied to A. Chinensis.
Other articles:
Literature Cited:
Rydberg (1916, p. 283) proposed Artemisia Vaseyana ...
Literature Cited:
Beetle (1959, p. 83) placed A. vaseyana as a subspecies of A. tridentata ...
Other articles:
Other articles:
Literature Cited:
Literature Cited:
From Nuttall's (1834a) catalogue of plants collected by Nathaniel Wyeth. Literature Cited:
Literature Cited:
Other articles:
Elliott (1823) published the name Brickellia in his Sketch of the Botany of South Carolina and Georgia.
Literature Cited:
Schilling, et al., 2015, infrageneric classification for Brickellia is behind a paywall, but available through Sci-Hub.
Species: — Brickellia cordifolia Elliott; … B. grandiflora Nutt.; …
…
Morphological trends:—Perennial herbs or subshrubs; leaves mostly opposite, mostly ovate, cordate to truncate,
larger leaves with naked basal veins, the lateral veins marginal at the point of divergence from the base of the midrib,
mostly petiolate; heads nodding (except for B. cordifolia and B. jaliscensis; these two species are also the exceptions
to the correlation of species from pine-oak forests having nodding heads); heads with numerous flowers, 25+, except
for B. jaliscensis 10–20.
…
5. Brickellia sect. Kuhnia (L.) E.E. Schill. & R. Scott, stat. nov. Kuhnia Linnaeus (1763: 1661).
Type: — Kuhnia eupatorioides L. [= Brickellia eupatorioides (L.) Shinners].
Species: — … ; B. eupatorioides (L.) Shinners; …
Morphological trends: — Herbaceous perennials, leaves usually alternate, linear to broadly lanceolate, involucre
strongly imbricate, bracts apices often densely glandular, heads with 8–12 flowers, pappus bristles usually plumose,
but barbellate in B. scoparia.
Taxonomic issues: — The wide-ranging B. eupatorioides is quite variable morphologically, and the status of its
named varieties, notably B. eupatorioides var. floridana (B. mosieri) needs to be evaluated.
9. Brickellia sect. Xerobrickellia Robinson (1917: 31).
Lectotype (here chosen): — Brickellia longifolia S. Watson.
Species: — … ; B. californica A. Gray; …
Morphological trends:
—
The leaves are petiolate, but the species of drier and lower elevations have shorter petioles.
The leaves ovate-rhomboid, becoming narrower and smaller in species of drier and lower regions; larger leaves have
the first pair of lateral veins diverging from the midrib above the leaf margin.
Flower number per head is 50–60 in B. cardiophylla and B. rhomboidea and 28–30 in B. brandegeei,
but fewer (13–ca. 25) in most other species (3–7 in B. longifolia; 8–12 in B. desertorum)
that are often found in somewhat lower and drier habitats.
Pendulous heads arose twice: once in the common ancestor of B. rusbyi and B. floribunda and,
again, in B. baccharidea and B. californica.
Taxonomic issues:
—
The widespread and variable B. californica and its relationship with B. desertorum remain in need of study.
Literature Cited:
Schilling, et al., 2015a, also behind a paywall, results supported the hypothesis that Brickellia is monophyletic and showed Barroetea, Phanerostylis, and Kuhnia all embedded within the genus.
Literature Cited:
Nesom (2022) revised Brickellia section Kuhnia, replacing our B. eupatorioides with B. suaveolens and B. leptophylla.
Other articles:
Literature Cited:
Other articles:
Locations:
Mora River.
Gray (1849) describes B. californica in his Plantae Fendlerianae Novi-Mexicanae.
The undescribed Mexican species is Brickellia wislizeni A.Gray, an accepted species that
occurs only in Mexico.
The dagger (†) refers to an undescribed Stevia in Coulter's Mexican collection.
Literature Cited:
Unfortunately, B. californica is not among the taxa studied by Schilling, et al., 2015.
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Linnaeus (1763) publication of Kuhnia eupatorioides from a collection in Pensylvania by Adam Kuhn.
Immediately above this entry is one for Ellisia nyctelea.
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Hooker (1840) described a single spcimen collected by Douglas.
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Nuttall (1841) described Brickellia grandiflora from collections on his trip with the Wyeth expedition,
citing Eupatorium grandiflorum Hook. as a synonym.
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Lamarck (1783, t. 1, p. 675-676) ...
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Linnaeus (1753, v. 2, p. 914) ...
Stśbe incana, cyano fimilis, tenuifolia. Bauh. pin. 273.
Stśbe auftriaca humilis. Cluf. hift. 2. p. 10.
Habitat in Auftria.
The genus name Stśbe now applies to a mainly south African Gnaphalieae.
At the time this was published, Austria included all of Hungary, Transylvania, Galicia, Bohemia, and Lombardy.
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Lamarck (1783, t. 1, p. 669) published C. maculosa ...
135. Ochsmann, Jorg. 2001. On the taxonomy of spotted knapweed (Centaurea stoebe L.). In: Smith, Lincoln, ed. Proceedings, 1st international knapweed symposium of the 21st century; 2001 March 15-16; Coeur d'Alene, ID. Albany, CA: U.S. Department of Agriculture, Agricultural Research Service: 33-41. [37829]
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Treated by Weber & Wittmann (2012) as Acosta maculosa (Lamarck) Holub.
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Treated by Ackerfield (2015) as C. stoebe ssp. micranthos (S. G. Gmelin ex Gugler) Hayek.
CABI.org (2021) says this ...
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FNANM (2021) says this ...
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Plants of the World (2021)does not accept subsp. micranthos (Gugler) Hayek.
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Twenty species of Cirsium “Thistle” are found in Colorado (Ackerfield, 2015). Eighteen of them are native, while two are introduced: C. arvense and C. vulgare. There are two endemic thistles: C. osterhoutii and C. perplexans.
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Wavyleaf thistle is listed by California as a noxious weed.
However, most reports of Cirsium undulatum in California are based upon misidentifications of C. canescens.
Cirsium undulatum is known to hybridize with C. flodmanii, C. hookerianum,
and C. scariosum var. coloradense.
J. T. Howell (1960b) reported that C. undulatum was suspected to hybridize with C. brevifolium in the Pacific Northwest.
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The genus Cirsium was first proposed by Miller (1754) in his Gardener's Dictionary.
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Nuttall (1818) published his Carduus undulatus in his Genera of North American Plants,
in the section Cnicus because the pappus is plumose.
He saw the plant on Lake Huron and in the upper Louisiana Territory.
I have not found a type specimen, but saw a note that the material might be in the Gray Herbarium.
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Frankton and Moore (1961) compare and contrast of Cirsium undulatum and C. flodmanii, both of which are known to occur in Colorado is behind a paywall, so I have reproduced the abstract here.
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Haffner and Hellwig (1999) investigated relationships within tribe Cardueae using ITS sequence data. The sole North American Cirsium in their study was C. texanum which does not occur in Colorado.
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Garcia-Jacas, et al. (2002) worked on tribal and subtribal delimitation of the Cardueae, unfortunately without any North American specimens.
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Kelch and Baldwin (2003) studied historical biogeography and ecology of true thistles (Cirsium, Cardueae, Compositae) in the New World.
Unfortunately, their sample of thistles did not include Cirsium undulatum.
However, three other native thistles, C. muticum, C. discolor, and C. rhaphilepis,
that commonly cluster with C. undulatum and did in fact cluster together in this study.
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Nuttall (1821, v. 2, p. 114) described a plant he found in the Arkansa Territory and introduced into the gardens of Philadelphia.
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Nuttall (1834a, v. 7, p. 29) described Crepis occidentalis
from a collection by Nathaniel Wyeth in 1833 between the Falls of the Columbia and the first navigable waters of the Missouri.
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In his Descriptions of new Species and Genera of Plants … Nuttall (1841) proposed Psilochenia occidentalis for his
previously published Crepis occidentalis Nutt.
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Enke (2009) is one of several papers by Enke and others exploring the genomics of the genus Crepis.
Unfortunately their data includes only one representative of the indigenous North American Crepis
— C. acuminata — which, I think, resulted in their results being somewhat unclear
with regard to that group.
Nevertheless, their data supports the maintenance of section Psilochenia within Crepis and
does not rule out elevation of the section to the rank of genus as Weber (1983) has proposed.
Note. — The North American species of Crepis sect. Psilochaenia are polyploid
and their placement within the genus remains unlcear at present.
*C. acuminata Nutt.
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Weber & Wittmann (2012) place the indigenous North American Crepis in Psilochenia also spelled Psilochaenia saying,
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Ackerfield retains our Largeflower Hawksbeard in Crepis as C. occidentalis Nutt.
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Nuttall (1818, v. 2, p. 185) described his Iva xanthiifolia from a collection made near Fort Mandan.
Fort Mandan was the name of the encampment which the Lewis and Clark Expedition built for wintering over in 1804-1805.
The encampment was located on now-private land on the Missouri River approximately twelve miles from the site of present-day Washburn, North Dakota, which developed later.
The exact location is unknown and may be partially submerged by the river.
There is a collection of Iva xanthiifolia attributed to Nuttall with a location of Upper Louisiana at the Harvard University Herbaria,
though there is no image and the type status is not indicated in the data record.
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Georg Fresenius is the apparent author of Index Seminum. Frankfort am Main, a list of seeds available from the Senckenberg Herbarium in Frankfort am Main.
In the 1836 Index, seeds are available for Cyclachaena xanthiifolia Fresen.
Footnote 4 in that document proposes Cyclachaena as a new genus.
However, there is no reference to Iva xanthiifolia Nutt. As the basionym.
Although no basionym reference was given Art. 41.4 (Melbourne Code) applies,
— International Code of Nomenclature for algae, fungi, and plants, 2012
— Melbourne Code text: © 2012, IAPT
— web-edition: © 2014, Paul van Rijckevorsel (all rights reserved)
Basionym
Iva xanthiifolia Nutt., Gen. N. Amer. Pl. [Nuttall]. 2: 185 (1818).
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Results support the dismemberment of Iva s.l. and recognition of Iva s. str., largely because Iva s. l. is shown to be paraphyletic. Most members of the section Cyclachaena were found to have relatively close interspecific relationships. Nevertheless cpDNA data strongly support two lineages within Cyclachaena. One lineage, including I. xanthiifolia, I. acerosa, I. nevadensis, and I. dealbata, has a close relationship with genus Euphrosyne.
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The Global Composite Checklist treats Iva xanthiifolia as an accepted name, and Cyclachaena xanthiifolia (Nutt.) Fresen. as a synonym. Plants of the World (Kew) does not follow their lead and accepts Cyclachaena xanthiifolia (Nutt.) Fresen..
Sometimes placed in Aster, Dieteria, or Macheranthera.
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Nuttall (1841, p. 300) proposed a new genus Dieteria without identifying a
type, the concept of which was lacking at the time.
I suspect that D. canescens (syn: Aster canescens Pursh)
is probably the closest thing we have to a type.
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Nuttall(1840, p. 300) in describing his plants from trip to the Oregon Territory, described a new genus for a subset of Asters.
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Locations:
Sandia Mountains.
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Gray, in Torrey (1857, p. [97] 41)
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Pursh (1814, v. 2., p. 547) ... Aster canescens ...
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Nuttall (1840, p. 300) published Dieteria and placed Aster canescens Pursh in it.
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Jacques Philippe Martin Cels (1740–1806) was a French botanist specializing in horticulture.
He started a botanical garden in which he cultivated foreign plants for sale, contributing to the growing public appetite for exotic flowers. He received and acclimatized numerous North American plants brought back by André Michaux and Louis-Augustin Bosc d'Antic. He strove to introduce many exotic species into France. The species in his garden were described by the botanist Étienne Pierre Ventenat (1757–1808) and illustrated by Pierre-Joseph Redouté (1759–1840) in Description des plantes nouvelles et peu connues, cultivées dans le jardin de J.-M. Cels, published in Paris in 1799 (WikiPedia, 2021).
Ventenat's (1801) description of Tagetes papposa states that it was discovered by A. Michaux in the countryside of Illinois.
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Hitchcock (1891, p. 503) moved our plant to Dyssodia papposa
in a catalogue of plants in Ames, Iowa.
Dysodia papposa, (Vent.)
—
Tagetes papposa, Vent. Hort. Cels. 1800.
—
D. chrysanthemoides, Lag. Nov. Gen. & Spec. 1816.
Sterile soil; frequent.
The pappus consists of “scales dissected into britles”
instead of “capillary bristles,” as it may at first appear.
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See my page about Ericameria nauseosa var. graveolens or “How did rubber rabbitbrush get that long scientific name?”
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Hall & Clements, 1923
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This key began as an abstraction from the Flora of North America (Urbatsch, et al., 2006) key to Ericameria parryi.
It was subsequently modified to be compatible with the keys in Flora of Colorado (Ackerfield, 2022)
and Colorado Flora, East Slope (Weber & Wittmann, 2012).
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Gray (1863, p. 66) ...
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Greene (1895c, p. 113) ...
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Nelson (1899, p. 373) ...
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Abstract
The genus Chrysothamnus (Asteraceae) contains 16 species. Recently, 4 species were transferred to Ericameria, and the remaining 12 were left in Chrysothamnus. The remaining species are now transferred to Ericameria as E. albida, E. depressa, E. eremobia, E. graminea, E. filfolia (formerly C. greenei), E. humilis, E. linifolia, E. molesta, E. pulchella, E. pulchelloides (a fossil species), E. spathulata, E. vaseyi, and E. viscidiflora. Section alignments are given, and some infraspecific combinations are also made.
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Infraspecific taxa accepted by Urbatsch, et al. (2006) are:
Taxa that are generally agreed to be found in Colorado are shown in bold face.
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Nelson (1899, p. 374) ...
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Nelson (1899, p. 374) ...
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Gray (1865, p. 541) published ...
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Greene (1895c, p. 113-114) ...
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Nelson (1899, p. 373) ...
Greene (1895c, p. 114) ...
Ackerfield (2022, p. 158) treats C. parryi (A. Gray) Greene ssp. nevadensis (A. Gray) Kittell as a synonym of
E. parryi var. attenuata (M. E. Jones) G. L. Nesom &G. I. Baird.
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This is the autonym of the plant described by Gray (1863).
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Usually looked upon as an ubituitous weed.
However, some sources, such as Plants of the World (Kew, 2026) suggest it is native to temperate North America,
and a good part of South America.
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Linnaeus (1753, vol. 2, p. 863) ...
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The Erigerons that are known from collections in Golden s.l. are shown below, and are placed in taxonomic groups of Nesom (2008).
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There are two collections of “Cutleaf Daisy” — Erigeron compositus Pursh —
made on the higher edges of Golden s.l. of Lookout Mountain.
This is consistent with other collections made in Jefferson County, and throughout Colorado,
with a few collections made on the higher plains, such as the Palmer Divide,
but generally in the foothills and higher mountain ranges.
The species was described by Pursh (1814) from Lewis & Clark collections on the Kooskoosky (Clearwater) River.
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Locations:
Kooskoosky River.
Of the western Erigerons found in Golden s.l. the first to be described was E. compositus Pursh.
The holotype was collected by Meriwether Lewis on the Kooskoosky (Clearwater) River, date unknown, and the voucher is now at ANS.
Pursh (1814) described it as follows:
My collection of this taxon was made as Little Scraggy Peak, (Buffalo Creek Recreation Area), Jefferson County, Colorado.
A. Gray (1862) proposed variety discoideus. The name is now treated as a synonym of the species.
Macoun (1884) proposed variety glabratus, but the name is not now recognized. The name is now treated as a synonym of the species.
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The first publication of the entity we now call Erigeron divergens was by Nuttall (1840).
It might be helpful to remember that the plains of Oregon would include a good part of southwest Wyoming.
The name E. divericatum had been previously used by Michaux (1803) and was not available for use by Nuttall.
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Torrey & Gray (1841, p. 175) in Flora of North America
proposed Erigeron divergens for the small fleabane that
was previously illegimately published by Nuttall as E. divaricatuum.
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Locations:
Santa Fe River.
Gray (1849, p. 68) described E. flagellare from a Fendler collection along Santa Fe Creek.
The “preceding species” and “foregoing” was Erigeron cinereum (sp. nov.),
now treated as a synonym of E. tracyi Greene.
The head was described as “… as large as those on Bellis perennis …”
and the involucre was described as “… hirsutum duplo superantibus …”
[… coarse erect or ascending hairs doubly … ??? ]
The asterisk “*” refers to some collections sent by Mr. Spaulding from Oregon,
which I don't think are particularly relevant to E. flagellaris.
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Nuttall (1818) described E. pumilum found on the plains of the Missouri.
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The first of the Erigeron found in the Golden, Colorado area was described from a collection made in Pensylvania.
Willdenow (1803) published the name from a description written by Mühlenberg.
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DeCandolle (1836) proposed Stenactis strigosa DC.
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Stenactis is currently treated as a section of Erigeron (Nesom, 2008).
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Some of the current synonyms for E. tracyi are:
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The first known description of the taxon we now call E. tracyi was by A. Gray (1849) in Plantae Fendlerianae Novi-Mexicanae, a description of plants collected by Augustus Fendler in New Mexico in 1841
Unfortunately, the name was unavailable, having been previously used by Hooker and Arnold for their E. cinereus, a native of Chile.
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Gray (1852, pt. 1, p. 91) places his E. cinereum as a variety of E. divergens in Plantae Wrightinae.
He make no comments that describe his thinking.
Gray (1852, pt. 2 p. 77) also places his E. cinereum as a variety of E. divergens in Plantae Wrightinae, part 2.
This placement is still accepted by Harrington one hundred years later (Harrington, 1954, 1964 2nd ed.).
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These plants were collected by the author while engaged in the State Geological Survey of Texas, during 1860 and '61.
Specimens of them are in the herbarium of the Academy of Natural Sciences at Philadelphia,
and also in the herbarium of Elias Durand, Esq.
…
Erigeron (Eurigera) nudiflorum, s. n.
—
Hirsutum pumilum, ramosum;
foliis lineari-oblongis, integris, sentis, confertissimis;
ramis apice longe nudis, monocephalis;
ligulis albis, plurimis, subuniseriatis, involucrum duplo excidentibus, acheniis oblongis glabris vel parum pilosis;
pappo radii et disci conformi, duplici;
exteriore breve setacea.
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E. L. Greene (1902) then published two names for what we now treat as a single species.
The names he published were: Erigeron commixtus and E. tracyi.
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E. commixtus was published before E. tracyi, quite literally a matter of a few inches.
It would therefore seem to have priority over E. tracyi.
Samuel Mills Tracy (1847-1920) was the first Director of the Mississippi Experiment Station.
Tracy is well known for his work in the taxonomy of various grasses, plant breeds,
and for adapting numerous forage plants to the biomes of the Southern United States.
He is perhaps best known for his work, Flora of Missouri, which was published in 1886.
Franklin Sumner Earle (1856-1929), served as the superintendent of the Mississippi Agriculture Experiment Station (1892–1895).
Soon after that Earle worked as a biologist and horticulturist of the Alabama Agriculture Experiment Station (1895–1900).
Tracy and Earle to not appear to be part of any particular expedition.
Perhaps, they were just two old botanists out for some fun field work.
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It is interesting to me that A. Nelson does not mention Greene's (1902) publication of Erigeron tracyi, whereas other names of Greene are acknowledged.
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Tidestrom (1925) recognizes Buckley's E. nudiflorus.
The author also recognizes E. divergens Torr. & Gray, and E. flagellaris Gray.
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Tidestrom and Kittell (1941) recognize Buckley's E. nudiflorus.
The authors also recognize E. divergens Torr. & Gray, and E. flagellaris Gray.
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Cronquist (1947) accepts Grays (1852) treatment of E. tracyi as a variety of E. divergens.
Along the way his key uses arrangement of hairs on stems as as a character to distinguish between E. divergens and E. flagelaris:
The variety cinereus is what we are calling E. tracyi today.
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Harrington (2nd ed., 1964, p. 565) placed E. tracyi as a variety of E. divergens.
This repeats his treatment of the taxon in his first edition (Harrington, 1954).
Of note is Harrington's acceptance of E. commixtus but not E. tracyi.
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Nesom (1989c) accepted E. tracyi as having priority over E. colomexicanus.
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Volume 20 of Flora of North America North of Mexico was published in 2006.
The treatment of Erigeron was written by Guy L. Nesom.
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Ackerfield's (2015) description of E. tracyi does not describe the stem hairs, or the direction in which they point.
Also, I think the stated length of the stoloniferous branches is quite short.
Perhaps, the length should be expressed in decimeters, such as “... stoloniferous branches, 1-3(6) dm. Long.”
Finally, the only synonym identified is E. colomexicanus perhaps because it has long been used in Colorado and New Mexico, but
E. commixtus and E. nudiflorus are not noted.
Couplet 24 of Ackerfield's key to Erigeron addresses the question of stem hairs:
Couplet 25 eventually ends up at Couplet 29 which differentiates between E. tracyi and E. divergens.
Couplet 34 eventually leads to E. flagellaris.
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Pursh (1814, v. 2, p. 573) ...
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Cavanilles (1791, v. 3, p. 41),
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Moore, et al. (2012) used DNA sequence data from the nuclear ribosomal ITS and ETS and chloroplast
psaI-accD regions to revisit hypotheses on biogeographic history across the genus.
Grindelia as a whole is well-supported
and is composed of two sister clades, one native to South America and the other native to North America including Mexico.
The North American taxa constitute two clades that largely occur on different sides of the Continental Divide.
The diverse radiation of Grindelia in the California Floristic Province appears to be most closely related to species from the Great Basin
and Colorado Plateau and evidently descended from drought-adapted ancestors.
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Greene, E. L.
1896.
Pittonia. v. 3, part 15.
New or Noteworthy Species, XVII pp. 91-116
pp. 102-103
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Rydberg (1906, p. 338) accepts G. inornata.
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Harrington (1964, 2nd ed.) accepts G. inornata.
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Strother & Wetter (2006), writing in Flora of North America treat G. inornata as a synonym of G. hirsutula Hook. & Arn.
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Weber & Wittmann (2012) treat G. inornata as a synonym of G. hirsutula.
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Ackerfield (2022) accepts G. inornata as a Colorado endemic.
Nesom (2025) accepts G. inornata.
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Plants of the World Online (Kew, 2025) accepts G. inornata, whereas World Flora Online places it in synonomy with G. hirsutula Hook. & Arn.
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First published in Mag. Neuesten Entdeck. Gesammten Naturk. Ges. Naturf. Freunde Berlin 1: 259 (1807)
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Pursh (1814, v. 2, p. 559) ...
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Dunal (1819, t. 2, p. 50) …
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Greene (1898, v. 3, n. 18, p. 297) in Sect. 6. Miscellaneous New Species
describes G. subalpina.
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Pursh (1814, v. 2, p. 540) described Solidago Sarothrae from a Lewis and Clark collection.
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Lagasca (1816, p. 30) published Gutierrezia as a place to put something that had no place.
Char. gener. differ.
Anthodium imbricatum , squamis apice reflexis :
Corolla radiata.
Receptaculum favosum, dentato-paleaceum.
Pappus polyphyllus, paleaceus.
Planta suffruticosa, glabra, resinosa.
Folia linearía, acuta, integerrima, vix lineam lata ,
pollicaria et amplius, superiora breviorŕ,
sessilia.
Corymbus pauciflorús,terminalis.
Anthodium oblongum, 8-florum.
Radius triflorus ligulatus.
Hab. in N. H. ♃
Spanish botanist Mariano La Gasca y Segura officially established the plant genus Gutierrezia in 1816
but did not leave a specific record of exactly which Gutierrez he was honoring.
However, botanical historians and researchers (such as Kathleen H. Keeler) strongly conclude the namesake is Pedro Gutiérrez Bueno (1745–1826).
Pedro Gutiérrez Bueno was a highly prominent Spanish chemist, apothecary, and pharmacist.
During the early 19th century, Gutiérrez Bueno was a leading member of the scientific community in Madrid.
At the very same time, La Gasca served as the director of the Real Jardín Botánico de Madrid,
making it highly likely La Gasca named the genus to recognize his colleague's contributions to science.
(Google AI, 4 June 2026).
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Nuttall (1818, v. 2, p. 163) published a new genus Brachyris and the specific name B. Euthamiae placing Pursh's (1814) in synonomy.
As it turns out, though, Lagasca's genus name Gutierrezia and Pursh's specific name sarothrae will have priority.
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Torrey & Gray (1838-1843, v. 2, p. 193) recognized that our plant belonged in Gutierrezia.
However, they adopted Nuttall's (1818) specific name Euthamiae
when Pursh's (1814) Sarothrae had prioirty.
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Britton & Rusby (1887, p. 10) placed our plant in Gutierrezia and applied Pursh's (1814) specific name sarothrae.
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As it happens, Lane (1982) designated a neotype for
Gutierrezia linearifolia:
MEXICO BAJA CALIFORNIA [Norte]:
1 mi. S Las Juntas (Sierra Juarez), Moran #16563, 6 Sep 1969
(Neotype: LL; isotype: SD).
Lane also placed G. linearifolia Lag. in
synonomy under G. sarothrae (Pursh) Britton & Rusby (Lane, 1980).
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Linnaeus (1753, v. 2, p. 904) ...
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Torrey & Gray (1842, v. 2, pp. 324-5) published H. nuttallii though apparently Nuttall, himself, had placed the name H. californicus on
specimens in the herbarium of the Academy of Arts and Sciences in Philadelphia.
Thomas Nuttall collected the type on the “plains of the Lewis River” – the Columbia River – and
intended to give it the name of Helianthus californicus.
However, the name was unavailable having been previously used by DeCandolle (1836) for a collection in California by David Douglas.
Torrey & Gray (1842) applied the name H. nuttallii giving credit to Nuttall for the collection.
An image of the type may be available at JSTOR (https://plants.jstor.org/stable/viewer/10.5555/al.ap.specimen.ph00014119).
However, it is securely locked away behind a paywall.
Nuttall (1840-1841) does not mention this in his description of Compositae from his 1834-5 expedition across the continent.
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The synonyms (POWO, 2022) of H. pauciflorus Nutt. ssp. pauciflorus,
which does not occur in Colorado, in order of date published, are:
Synonyms (POWO, 2022) of H. pauciflorus Nutt. Ssp. subrhomboideus (Rydb.) O. Spring & E. Schilling,
which does occur in Colorado, again in order of pub;ication date, are:
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On his 1811 trip up the Missouri River, Thomas Nuttall collected a sunflower that has become our Helianthus pauciflorus.
The location is given as Lower Louisiana.
Since the current range of our plant is Missouri and northward,
one would assume the collection was not made south of Missouri.
He apparently brought seeds of this plant back to England.
I have seen references to plants growing in Lambert's garden, the Chelsea garden, and the royal garden of Paris.
These garden-grown plants can be found in the text below and would gave rise to multiple names for Nuttall's plant.
I have not seen any reference to collected plant material, either in US herbaria, or the British Museum.
Nuttall (1818, v. 2, p. 177) published this plant on April 3, 1818, as Helianthus pauciflorus.
The actual date of publication is important because there were two more names published later in 1818; those by Cassini (1818) and Sims (1818)
Literature Cited:
Cassini (1818) published the same plant grown from seeds in the Jardin du Roi of Paris.
This name was published in September, 1818, per Kartesz & Gandhi (1990), below.
Literature Cited:
Meanwhile Sims (1818) Curtis's Botanical Magazine, Plate 2020 and following discussion, published a third name for our plant using an image from Lambert's garden and a description from the Botanic Garden at Chelsea.
This name was published in October, 1818, per Kartesz & Gandhi (1990), below.
From its rambling manner of growth, this plant does not
appear very sightly in the garden ; but is one of the most
desirable flowers imaginable for ornamenting rooms, as it
lives long in water, and from its spreading branches, the
flowers dispose well, and make a very brilliant appearance.
A hardy perennial. Blooms in August and September.
Our drawing was made from a specimen communicated by
Aylmer Bourke Lambert, Esq. three years ago, out of his
collection at Boyton, where it was raised from seeds collected
on the borders of the Missouri, by Mr. Nuttall ; but our
description was taken at the Botanic Garden at Chelsea in
August last.
Literature Cited:
Other articles:
Nuttall's expedition to Arkansas spanned 1818 to 1820, though he was in that state mostly in 1819.
There is a voucher in the British Museum (BM001025610) that appears to be from Nuttall's trip to Arkansas.
It is currently filed as Helianthus rigidus (Cass.) Desf. subsp. rigidus (Cass.) Desf.
Nuttall (1834b) does not mention any Asteraceae in his "Collections towards a flora of the territory of Arkansas."
He will though in Nuttall (1841) "Descriptions of new Species and Genera of Plants. "
Literature Cited:
Cuvier (1821, v. 20, p. 300) in his dictionary of natural sciences repeats Cassini's (1818) diagnosis and name,
while noting that Rene Dessfontaine (at the Jardin du Roi) thinks Sim's (1818) H. diffusus may be the same plant.
“H. Cass.” = Count Alexandre Henri Gabriel de Cassini (9 May 1781 – 23 April 1832) was a French botanist and naturalist, who specialised in the sunflower family (Asteraceae) (then known as family Compositae).
Literature Cited:
Desfontaines (1829) in his 3rd edition of the catalog,
in English, “Catalog of the Plants of the Royal Garden of Paris with annotations on new or less known plants”
published Helianthus rigidus, placing H. diffusus and Harpalium rigidum in synonomy.
Literature Cited:
Other articles:
Nuttall's (1841) "Descriptions of new species and genera of plants in the natural order of the Compositae" was mostly about his 1834 travels across the continent, though he also published plants from the Arkansas trip and some of those collected by William Gambel.
He also published Helianthus crassifolius, now treated as a synonym of Helianthus pauciflorus ssp. pauciflorus.
Literature Cited:
Torrey & Gray (1841, v. 2, p. 323-331) Flora of North America accepts H. rigidus Desf.
and treats H. pauciflorus Nutt. as an obscure or little-known species.
I would speculate this was the result of no actual plant material being available in America.
β
branches or peduncles simple, elongated; scales of the involucre
ovate-lanceolate or ovate, more acute.
—
H. diffusus, Sims, bot. mag. T. 2020 (poor.)
H. Missuricus, "Spreng. Pug. P. 21 ;"
Link. Enum. 2. p. 352.?'
H. Missouriensis, Schwein. ! Herb. ;
Nutt. ! In trans. Amer. Phil. Soc. I. c.
H. atrorubens, Bot. reg. t. 508 ;
Hook. ! Fl. Bor.-Am. 1. p. 312 ;
DC. Prodr. I. c. ex char. ; not of Linn.
Plains and prairies, western part of Georgia, Elliott !
Illinois, Mr. Buckley ! And St. Louis, Drummond! To the Upper Missouri, Nuttall!
Mr. Nicollet! Arkansas, Nuttall! Louisiana, Dr. Leavenworth! And Texas,
Drummond!
β Missouri, Nuttall! (spec, cult.) and Saskatchawan, Drummond!
Aug.-Sept.
—
Plant stout, 1-3 feet high, rather naked above ; the
rigid stem hispid with short papillose hairs, often smoothish above : the
coriaceous thick leaves (cauline 3-5 inches long, an inch or less wide) very
rough with short bristles arising from papillas, which give the surface a
cinereous hue, and often a whitish blistered appearance.
Heads few, but showy ;
the disk about an inch in diameter; the 20-24 rays an inch long. Scales of
the involucre regularly and closely imbricated in 3-4 series. Receptacle
convex : the chaff entire, ciliate on the back towards the summit. Lobes of
the disk-corolla purple : style yellow. Achenia somewhat lenticular, hirsute
when young, but more smooth when mature, except the edges and summit.
The pappus is variable, even in different flowers from the same individual ;
consisting either of the two scale-like awns without intermediate squamellae,
or of 1-4 of the latter on each side, or with the squamaellae confluent
with the lateral scales, which become dilated and auriculate or lobed
at the base, &c. &c. ; whence we conclude that Harpalium, Cass. Is
founded upon insufficient and very inconstant characters.
The var. β differs but slightly,
and passes completely into the other forms of this well-marked species,
so that it hardly merits to be distinguished. In some of the
Texan specimens, and in those described by Nuttall as H. crassifolius, the
leaves are more attenuated to each end, more serrated, and the upper surface
comparatively smooth. — The stem sometimes exudes resin in small quantity.
Lower Louisiana. — Plant 4-5 feet high. Leaves sometimes ternately
verticillate, very long, paler beneath and somewhat pubescent. Ray and
disk nearly the same color. Nutt. — This species is unknown to us ; we find
no specimen in the herbarium of the Academy of Natural Sciences.
Literature Cited:
Porter & Coulter (1874, p. 71) accepted Helianthis rigidus Desf.
I don't see any vouchers in SEINet determined Helianthus pauciflorus or H. rigidus from Colorado
that pre-dates 1874 publication date.
Nor do I see any Hall & Harbour or Brandegee collections of either taxon.
These searches in SEINet would include the Gray Herbarium and New York Botanic Garden by default.
Literature Cited:
Gray (1884, v. 1, pt. 2, p. 274-175) accepts H. rigidus Desf. as
H. pauciflorus Nutt. slips farther into obscurity.
“ *** Stem and (mostly opposite and triplinerves) leaves more or less hispid,
hirsute, or scabrous ... :
chaff of receptacle enture or some 3-toothed at the apex, pointless :
rays numerous and conspicuous”
“ ** Disk of the head dark purple”
Literature Cited:
Coulter (1885, p. 186) also accepted Helianthis rigidus Desf. but gave no sources.
Literature Cited:
Rydberg (1900, p. 419) in his Catalogue of the Flora of Montana and Yellowstone National Park publishes H. subrhomboideus Rydb.
Literature Cited:
Rydberg (1906, p. 373) accepted H. subrhomboideus replacing H. rigidus in part.
I don’t think ne intended to say that A. Gray was the author of H. rigidus.
Instead I think he meant that some of the plants that A. Gray would describe as H. rigidus should be segregated into H. subrhomboideus.
Literature Cited:
Coulter & Nelson (1909, p. 548) accept H. scaberrimus Ell., placing H. rigidus Desf.
and H. subrhomboideus Rydb. In synonomy.
Literature Cited:
Kartesz & Ganghi (1990) accept Helianthus pauciflorus Nutt.
Kartesz & Gandhi (1990b) accept Helianthus pauciflorus Nutt. ssp. subrhomboideus
(Rydb.) Spring & Schilling.
Literature Cited:
Spring & Schiling (1990) ...
Literature Cited:
Other articles:
Nuttall (1841, p. 366) described the Little Sunflower from plants he collected on plains of the Platte enroute to Oregon Territory.
Other articles:
The International Plant Names Index continues to use H. multiflorus, whereas nearly all other floras and checklists use the corrected Latin gender of H. multiflora.
Literature Cited:
Nuttall (1848, p. 171) described the genus Heliomeris and the species H. multiflorus from a collection by William Gambel in Upper California.
Literature Cited:
Literature Cited:
Other articles:
Literature Cited:
Other articles:
Nuttall (1840) described this from the Rocky Mountain plains,
near the banks of the Platte.
Heterotheca foliosa (Nutt.) Shinners was published in a journal called Field & Laboratory, contributions from the Science Departments of Southern Methodist University (Field & Lab. 19:71 (1951).)
Other articles:
Literature Cited:
Nuttall (1813) published Sideranthus integrifolius which is a nomen nudum.
Literature Cited:
Pursh (1814, v. 2, p. 564) ...
Literature Cited:
Publication of Heterotheca villosa by Shinners; not available online.
Other articles:
Other articles:
Literature Cited:
Locations:
Fort Carleton Provincial Historic Site.
Hooker, 1833, 1(6): p298
The Smoking River (often spelled Smoky River) is located in west-central Alberta, Canada. It originates in the Canadian Rockies near the continental divide and flows northeast before joining the Peace River. When Drummond reported finding Hieracium albiflorum (White Hawkweed) "north of the Smoking River in latitude 56°N" during his botanical explorations, he was referring to the remote wilderness region of northwestern Alberta, just east of the British Columbia border.
Other articles:
Literature Cited:
Gray, Asa. 1849. p. 114.
Literature Cited:
Schultz-Bipontinus, C. H. 1861.
Ueber die Hieracien Amerika's.
Bonplandia. Zeitschrift fu¨r die gesammte Botanik.
Vol. IX. No. 12. 1861.
pp. 172-175
https://www.biodiversitylibrary.org/page/28138259#page/181/
p. 173
C. H. Schultz-Bipontinus (born Carl Heinrich Schultz, 1805–1867) was a prominent German physician and botanist.
He specialized heavily in the sunflower family (Asteraceae).
He used "Bipontinus"—a Latinized reference to his birthplace of Zweibrücken (Two Bridges, or Bipons in Latin)—to distinguish himself from a contemporary botanist, Carl Heinrich Schultz-Schultzenstein (Google AI, 29 June 2026).
He was describing Hieracium (hawkweeds) from America for two primary reasons:
(1) he had the taxonomic expertise,
and (2) during the 19th-century expansion of botanical exploration,
American institutions and botanists (such as Asa Gray at Harvard) sent specimens back to Europe to be formally named.
Schultz-Bipontinus frequently published descriptions of these new American and Mexican species in European scientific journals,
most notably in the German botanical periodical Bonplandia.
Literature Cited:
Locations:
Beulah Valley.
Wet Mountain Valley.
Porter and Coulter, 1874, Synopsis of the flora of Colorado ...
Mace's Hole is now called Beulah Valley.
Literature Cited:
Subgenus Chioracium
Other articles:
Literature Cited:
Sprengel, Syst. Veg., ed. 16, 3:640 (1826)
Literature Cited:
A.Gray, Mem. Amer. Acad. Arts ser. 2, 4(1): 102 (1849).
Two pages later, i.e., p. 104, Gray will describe Amauria? dissecta, which we now know as Hymenothrix dissecta.
Other articles:
Literature Cited:
Locations:
Mora River.
Gray (1849, p. 104) was unsure what name to apply to Fendler's and Fremont's specimens because they were incomplete.
See also, Gray's Hymenothrix nov. gen., two pages previous.
Literature Cited:
Britton (1889, p. 68) published Bahia dissecta from collections by Dr. E. A. Mearns, in the Mongollon and San Francisco Mountains, Arizona.
Literature Cited:
Locations:
Mora River.
Rydberg (1915, v. 34, pt. 1 [1914]) published Amauriopsis with A. dissecta being the type species.
Flora of North America, Vol. 21, 2006 retains our plant in Amauriopsis.
Literature Cited:
Baldwin and Wood (2016) examined phyllotaxy within the Bahia alliance and, among other findings,
showed that merger of Amauriopsis and Hymenothrix into a common,
monophyletic genus within the Bahia alliance would simplify the taxonomy.
Hymenothrix has priority for such a group,
which is characterized
in part by alternate leaves, obtuse to acute style-branch apices,
often zygomorphic disc corollas (of outer florets), and pappus
scales generally ≥ 10 or absent.
Literature Cited:
Other articles:
Hooker (1833, 1(6), p. 317) published Hymenopappus filifolius
Other articles:
Literature Cited:
Osterhout (1918, p. 90) ...
Literature Cited:
Turner (1956, p. 237) ...
Other articles:
Other articles:
There is a single collection of Liatris ligulistylis (A. Nelson) K. Schum. “Northern Plains Gayfeather”
in Golden s.l. and indeed in Jefferson County.
State-wide, collections seem to be concentrated around Colorado Springs, and then scattered in odd locations in the interior mountains.
First described as Lacinaria ligulistylis A. Nelson Bot. Gaz. 31: 405. 1901,
it was placed in Liatris ligulistylis (A. Nelson) K. Schumann Just’s Bot. Jahresber. 29(1): 569. 1903.
Other articles:
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Superficially Lygodesmia juncea is very similar to Stephanomeria pauciflora.
However, it turns out they are quite distinct, being separated by consistent differences in cotyledon, achene, and pollen morphology
and base chromosome numbers (x = 9 in Lygodesmia; x = 8 in Stephanomeria).
Both Weber & Wittmann (2012) and Ackerfield (2015) separate them on morphology of the pappus, with Lygodesmia having a pappus of capillary brinstles
and Stephanomeria a pappus of plumose bristles.
Other articles:
Literature Cited:
Hooker.
Fl. Bor.-Amer. (Hooker) 2(7): 24 (1834).
Literature Cited:
Nuttall. 1841.
Descriptions of new species and genera of plants in the natural order of the Compositae :collected in a tour across the continent to the Pacific, a residence in Oregon, and a visit to the Sandwich Islands and upper California, during the years 1834 and 1835.
p. 390
Both Amida hirsuta Nutt. and A. gracilis Nutt. are now treated as synonyms of Madia glomerata Hook.
Other articles:
Literature Cited:
Pursh (1814, v. 2, p. 502) published it as Sonchus pulchellus
from an unknown dried specimen.
It was not a Lewis & Clark collection, per Moulton (1999).
Literature Cited:
Frequently referred to, but outside my price range, Bremer (1994) apparently made a strong case for treating our plant as Mulgedium pulchellum.
Literature Cited:
Flora of North America (2006), John Strother genus author, treats this taxon as Mulgedium pulchelllum (Pursh) G. Don
Literature Cited:
Weber & Wittmann (2012) treat this taxon as L. tatarica (L.) Meyer subsp. pulchella (Pursh) Stebbins.
Literature Cited:
Ackerfield (2015) treats this taxon as Lactuca tatarica (L.) C. A. Mey var. pulchella (Pursh) Breitung.
Literature Cited:
Plants of the World Online (Kew, 2021) treats it as Lactuca pulchella DC.
Other articles:
Literature Cited:
Pursh (1814, v. 2, suppl., p.742) published Troximon cuspidatum from a collection in
upper Louisiana by John Bradbury.
In Upper Louisiana.
Bradbury.
♃
v. s. in Herb. Bradbury.
Flowers large, yellow.
Resembles the preceding.
The preceding was Troximon glaucum (=Agoseris glauca)
Literature Cited:
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Nothocalais was first published by Gray (1884, p. 420) as a section of Microseris.
Literature Cited:
Greene (1886, p. 55) elevated Gray's section Nothocalais to the rank of species.
On bleak, stony hills and fertile prairies, from Dakota
and Colorado to Wisconsin and Illinois. Scarcely distinguishable
from its far Western congeners except by the
pappus. The undulate-crisped, white-hairy margins of the
grassy leaves of this giving it an aspect so strikingly unlike
the general appearance of the other species of his genus
Troximon, were points not overlooked by that well traveled
and most keenly observant botanist, Mr. Nuttall. That he
noticed the peculiarity and was impressed by it is evinced
by his effort to invest the species with a new specific name,
marginatum, more appropriate than Pursh's cuspidatum,
which was given to it in reference to the acuminate rather
than cuspidate bracts, and has, therefore, no fitness, but
which must needs be retained in deference to its priority.
The name marginatum would, indeed, be equally and in the
same way, applicable to each of the three known species of
Nothocalais.
Literature Cited:
Other articles:
Cassini (1817, p. 31) dscribed his new genus of Oligosporus
SEINet (2021+) data includes collections of Packera that have been
determined the following species:
Literature Cited:
Harrington (1964, 2nd ed., p. 610) distinguishes between Packera fendleri and P. plattensis by the hairiness of the achenes,
those of P. fendleri being glabrous, and those of P. plattensis hispidulous along the angles.
Literature Cited:
Love & Love (1976) published Packera
Literature Cited:
Weber & Love (1981) published fifty-one new names in Packera
Literature Cited:
Trock (2006) key to Packera has two ways to get into P. fendleri and four ways to get into P. plattensis.
Literature Cited:
Weber & Wittmann (2012) key focuses mainly on the shape of the basal leaves …
Ackerfield (2015) key also focuses on the shape of the basal leaves …
Other articles:
Locations:
Schweich Hill.
Fendler's Ragwort — Packera fendleri (A. Gray) W.A. Weber & Á. Löve —
is widespread and fairly common in dry meadows and slopes of Golden s.l..
The author has collected it on Schweich Hill,
and Deadman Gulch.
Others have collected the plant on North and South Table Mountains.
P. fendleri is frequently confused with P. plattensis which is more a plant of the great plains.
Literature Cited:
Other articles:
Locations:
Santa Fe River.
Gray (1849, p. 109) writing in Plantae Fendlerianae Novi-Mexicanae described P. fendleri from a
collection made “ ...along the Creek, twelve miles above Santa Fé …”
The current accepted name is Santa Fe River.
There follows a half page discussion of similar Senecio found throughout the southwest, such as Senecio multilobatus (Torr. & A. Gray, ined.)
Other articles:
Literature Cited:
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Other articles:
Locations:
Cottonwood Creek.
Rydberg (1900, v. 27, p. 175) ...
Literature Cited:
Weber & Love (1981, p. 48) published the name Packera tridenticulata.
Other articles:
Other articles:
Locations:
Santa Rita.
On the Peaks to Plains trail about 2/3 of the way from the Grant Terry Bridge to the Tough Cuss Bridge, look in the talus piles below the Canal Zone climbing area for a 4-5 foot shrub with clusters of small yellow flowers.
This is “Mountain Tail-Leaf” — Pericome caudatus A. Gray.
Though it is in the sunflower family (Asteraceae) it has only disk flowers, and not the large ray flowers that characterize the typical sunflower.
The leaves are triangular and have a long tail, giving it both its common name “Tail-Leaf” and scientific name caudatus, which means “having a tail.”
First collected by Charles Wright in 1851 who wrote that he found it “… among rocks, on the sides of mountains, at the copper mines, New Mexico …”
Mr. Wright was working with the scientific corps of the U. S. Boundary Commission.
Their itinerary from July to November 1851, was from El Paso to the copper mines of Santa Rita del Cobre in the southwestern part of New Mexico,
and then into the northern part of Mexico.
It seems certain that Wright’s collection of Mountain Tail-Leaf was made near Santa Rita del Cobre, about 12 miles east of Silver City, NM.
The town no longer exists, having been completely swallowed up by the Freeport-McMoRan Chino open-pit copper mining complex.
We now know our plant’s primary range is in the lower elevation mountains of Colorado, New Mexico, and Arizona, with a few scattered collections in northern Mexico, and eastern California. Near Golden, it is known only from Clear Creek Canyon, but also occurs around Pine and Buffalo Creek in central Jefferson County.
Other articles:
A. Gray (1853, p. 82) ...
Other articles:
Literature Cited:
Nuttall (1818, v. 2, p. 167-168) ...
Literature Cited:
Britton (1901, p. 1008) Flora of the Northern States and Canada ...
Other articles:
Other articles:
Literature Cited:
DeCandolle (1837, v. 6, p. 228) ...
Literature Cited:
Anderberg (1991) proposed moving G. canescens to Pseudognaphalium Kirp.
Other articles:
Literature Cited:
Greene (1902, v. 15, p. 278) published Gnaphalium maccounii
Literature Cited:
Rafinesque (1818, p. 268) in his review of Pursh (1814) Flora Americae Septentrionalis ...
Other articles:
“Upright Prairie Coneflower” Ratibida columnifera is predominantly a Great Plains species which
extends from southeastern British Columbia to Manitoba and Michigan,
south through Illinois to Louisiana, and west through Texas and northern Mexico to Arizona.
Literature Cited:
Other articles:
Fraser's (1813) catalogue of new and interesting plants published for the first time a number of new taxa for North America.
Many of these were Thomas Nuttall collections that were subsequently published in
Pursh (1814) Flora Americae Septentrionalis and
Nuttall (1818) Gen. Am.
75 * Rudbeckia columnifera.
‡
Spontaneous varieties of this plant sometimes occur with bright fulvous flowers,
colored like Tagetes patula :
the stem is simple, seldom producing more than three flowers,
which are of an uncommon length,
appearing like a column of flosculi,
subtended by 5-8 neutral florets,
and a simple calyx.
76 Rudbeckia purpurea.
* serotina.
‡
Stem somewhat branching and hirsute,
flowers brighter and more numerous.
Copies of Fraser's (1813) catalogue were quite rare, so Greene (1889) republished it.
It is unclear who actually wrote the catalogue.
Greene was convinced that Nuttall wrote it having seen an original copy sent to Zaccheus Collins from Messrs. Fraser, saying
Literature Cited:
Other articles:
Pursh (v. 2, 1816) published the name, citing Fraser's (1813) catalogue.
Literature Cited:
Other articles:
Nuttall (1818) published Rudbeckia columnaris
In this publication, Nuttal has altered R. columnifera to R. columnaris
not referred to R. columnifera as published in Fraser's (1813) Catalogue.
I suspect this might be enough to render R. columnaris and illegitimate name.
If so, this will have an impact on Rafinesque's proposal of Ratibida columnaris Raf., nextunder.
Literature Cited:
Other articles:
Rafinesque (1818) wrote a two-part review of Pursh (1814) Flora Americae Septentrionalis.
The reference to “Fl. Miss.“ is a puzzle. I have seen another reference to this publication that it was “ined.”
However, it is unclear whether “Miss.” refers to either Mississippi or Missouri.
Literature Cited:
Type locality: Upper Louisiana.
Range: British Columbia and Saskatchewan to Arizona, Texas, and Tennessee.
New Mexico: Sierra Grande; mountains west of Grants Station;
Santa Fe and Las Vegas mountains;
Clayton; Lower Plaza; White and Sacramento mountains.
Plains and low hills, in the Upper Sonoran and Transition zones.
Type locality: “In Mexici provinc. Texas as San-Fernando de Bejar, et in sinu Spiritus-Sancti ad lacum Sancti-Nicolai.”
Range: With the species, but more common in New Mexico.
New Mexico: Dulce; Chama; Pecos; Santa Antonita; Ramah;
near Las Vegas; mountains west of Grants Station; El Cedro;
Tucumcari; Mongollon Mountains; White Mountains; Buchanan;
Redlands; Queen; Knowles; Artesia.
Literature Cited:
Other articles:
Reveal (1968) summarized the evidence and opinions on the author of the names in Fraser's (1813) Catalogue
and the legitimacy of them. Excerpts from Reveal's introduction were presented above, whereas his specific comments
on Ratibida columnifera are below.
Literature Cited:
Monograph on Ratibida
Other articles:
Other articles:
Other articles:
Literature Cited:
Linnaeus (1753, vol. 2, p. 906) ...
Other articles:
Literature Cited:
Nelson (1901, p. 234-235) ...
Literature Cited:
In Vascular Plants of the Pacific Northwest (Hitchcock, et al., 1955, Part 5),
Cronquist outlines that while the western montane populations (originally named R. ampla) possess distinct physical dimensions,
they are not completely isolated morphologically from the highly variable eastern populations of R. laciniata.
The Key Distinctions: He notes that var. ampla primarily differs from the typical eastern variety (var. laciniata) by having significantly larger receptacles (the disk centers) and larger flower heads/disks altogether.
The Taxonomic Choice: Because the underlying structural features of the leaves, stems, and disk florets remain overwhelmingly identical, and because R. laciniata is widely recognized as a complex, variable species across North America, Cronquist argued that a varietal status was much more appropriate than keeping it separated as an independent species.
By publishing it as Rudbeckia laciniata L. var. ampla (A. Nelson) Cronquist,
he formalized the idea that it represents a robust, geographically distinct western expression
of a single, highly adaptable widespread species.
Literature Cited:
Other articles:
Locations:
Marshall Pass.
Greene, Pittonia 3(15): 105 (1896). proposed S. accedens ...
In open woods at Marshall Pass, Colorado, 4 Sept., 1896 ;
barely in flower at that date.
Plant with the foliage of S. cernuus nearly,
but of different mode of growth and inflorescence ;
the large nodding heads almost those of S. Rusbyi.
Other articles:
Literature Cited:
Senecio bigelovii A.Gray, Pacif. Railr. Rep. 4(5, 4): 111 (1857). ...
Gray described Senecio bigelovii in 1857;
when he subsequently described var. hallii in 1863,
the publication of the variety necessarily established the autonym S. bigelovii var. bigelovii.
Other articles:
Gray described the parent species, Senecio bigelovii, in 1857 (see above) and six years later described a hairy variety, S. bigelovii var. hallii.
Literature Cited:
Other articles:
Gray's (1863) enumeration of the plants collected by Hall & Harbour ... and, of course, Parry,
but he isn't mentioned.
Also note that Gray places S. megacephalus Nutt. in synonomy with S. bigelovii var. hallii.
However, Barkley (2006) accepts both names as valid taxa placing them in different groups.
See S. megacephalus, below. The taxon is found mostly in Idaho and Montana and is not known from Colorado.
Literature Cited:
Other articles:
Locations:
Marshall Pass.
Edward L. Greene (1896, p. 110) proposed Senecio accedens for a plant collected at Marshall Pass.
See text, above.
Rydberg (1917) will accept this name but mis-spell it.
Literature Cited:
Rydberg (1917, p. 987)
...
accepted Senecio accidens, mis-spelling S. accedens Greene (1896).
Rydberg placed S. bigelovii var. hallii A. Gray i synonomy.
Other articles:
Literature Cited:
Rydberg (1910, v. 37, p. 467-8) published S. ambrosioides as a primarily Colorado species
and S. kingii as a primarily Utah species.
Other articles:
Other articles:
coarctate = crowded together.
Literature Cited:
Cronquist. 1950. Notes on Compositae of Northwestern United States
made a new combination in Senecio ...
Gray (1863) placed S. megacephalus in synonomy with S. bigelovii var. hallii,
although Barkley (2006) treats them both a valid taxa, even in separate groups.
S. megacepahlus is found mostly in Idaho and Montana and is not known from Colorado.
Literature Cited:
Other articles:
Nuttall (1841, p. 410) described our plant ...
Other articles:
Other articles:
Locations:
Schweich Hill.
Known from Apex Park and Tin Cup Ridge, North and South Table Mountains, and Schweich Hill.
Literature Cited:
Other articles:
Locations:
Jeffrey City.
Senecio spartioides was described by Torrey and A. Gray (1843) from a
Lt. John Fremont collection.
Other articles:
Literature Cited:
Semple (2018) is a frequently updated resource on Goldenrods (Solidago sp.).
Other articles:
Other articles:
Aiton (1789, v. 3, p. 211) published Solidago gigantea from plants grown at the
Physick Garden at Chelsea.
Other articles:
Literature Cited:
Other articles:
Nuttall (1834a, p. 32-33) published Solidago missouriensis …
Other articles:
Literature Cited:
Other articles:
Nuttall's (1841) description of S. nana.
Other articles:
DeCandolle (1836, v. 5, p. 332) ...
Other articles:
Literature Cited:
Kunth (1818, t. 4(15), p. 81) ....
Other articles:
Literature Cited:
Other articles:
Other articles:
Literature Cited:
Gray (1876, p. 58-59) publisher S. sparsiflora from a collection on the Wheeler Expedition.
Other articles:
Other articles:
Other articles:
Literature Cited:
Torrey (1827, p. 210) published several of Dr. James collections,
placing them all tentatively in Prenanthes.
Literature Cited:
Coulter & Nelson (1909, p. 588) placed P. pauciflora Torr. in
Stephanomeria
Other articles:
Literature Cited:
Linnaeus (1753, v. 2, p. 875) ...
Flora Virginica, published as two volumes by Jan Frederik Gronovius in 1739 and 1743,
based on the work of colonial Virginia botanist John Clayton.
Flora Leydensus Prodromus, published by Adriaan Royen in 1740.
Royen was a professor of botany at Leyden and well aquainted with both Cliffort and Linnaeus.
Johan Jacob Dillenii, 1732. Hortus Elthamensis, a catalogue of the rare plants growing at Eltham, London.
Literature Cited:
Other articles:
Other articles:
Literature Cited:
Other articles:
Locations:
Canadian River.
Ocate Creek.
Plantae Fendleriana
Other articles:
Other articles:
Other articles:
Literature Cited:
Gray (1884, v. 1, pt. 2, p. 192) ...
Other articles:
Literature Cited:
Publication of Aster ericoides L. var strictus Porter in Porter and Coulter (1874, p. 56) ...
Literature Cited:
The basionym: Asteraceae Aster porteri A.Gray Proc. Amer. Acad. Arts xvi. (1880, p. 99) ...
Literature Cited:
Multivariate morphometric analyses were used to compare inflorescence and head traits of 313
individuals of Symphyotrichum subsect. Porteriani.
Other articles:
Other articles:
Other articles:
Literature Cited:
Sprengel (1826, v. 3, p. 454) ...
Literature Cited:
Kuntze (1898, Pt. 3[3], p. 182) ...
Plants of the World (Kew) does not accept Kuntze's authorship of this name,
instead accepting Thelesperma megapotamicum (Spreng.) Herter, Revista Sudamer. Bot. 7: 235 (1943).
I am unable to find that publication online.
Other articles:
Literature Cited:
Hooker (1834,v. 2. p. 16) described Townsendia and T. sericea
that will become a synonym of T. exscapa.
Other articles:
(Nuttall, 1840, p. 306) ...
Other articles:
Literature Cited:
Nuttall (1840-1841, p. 304-305) describes a collection as T. sericea
which is later to be determined T. hookeri
Literature Cited:
Beaman (1957, p. 95-100), in a six-page section described Townsendia hookeri
from an Ira W. Clokey collection in Mount Vernon Canyon.
This seems to be the only collection made by Ira W. Clokey on 14 April 1920,
indeed the only collection Mr. Clokey made in the week from 10 April to 16 April 1920.
Clokey also collected a large number of duplicates, i.e. 18 duplicates according to Beaman (1957).
SEINet reveals 3 more duplicates, at IND, NO, and MIN.
Other articles:
Other articles:
Literature Cited:
Nuttall (1813) may have been first into print with:
Unfortunately, Sideranthus was a nomen nudum when this was published,
as the name was not validly published until 1840.
Literature Cited:
Pursh (1814, v. 2, p. 564) ...
In open prairies on the Missouri.
M. Lewis. ♃. Aug. Sept.
v. s. in Herb. Lewis.
Rays yellow.
Tota planta rigida, scabra, cano-viridescens.
Caulis ramosissimus.
Rami angulosi.
Ramuli laterales breves, uniflori.
Folia alterna, bipinnatifida, rigida, laciniis linearibus, spinula terminatis, inferioribus inciso-dentatis.
Flores laterales solitarii et terminales congesti, subcorymbosi, lutei.
Calyx imbricatus : squamis exterioribus brevioribus, acutis, hirsutis ;
interioribus linearibus, scariosis.
Corollulae radii lineari-lanceolatae, bidentatae.
Receptaculum : Paleis brevibus, subuiatis.
Both species are an intermediate link between Aster and Inula.
They approach the nearest to Amellus ;
but the receptacle is not, properly speaking, paleaceous, but only setaceous.
Literature Cited:
Moulton (1999) lists the Lewis & Clark collection as Machaeranthera pinnatifida
Literature Cited:
Morgan & Hartman (2003, v. 20, p. 1406) ...
Date and time this article was prepared:
10/2/2026 12:51:48 PM
- Kunth, Carol Sigismund, 1815-1825.
2. VERONICA xalapensis. †
2. VERONICA xalapensis. nov. sp.
V. caule erecto , pubescente ; foliis oblongis , cernulatis ;
floribus axillaribus , solitariis , breviter pedunculatis , folio brevioribus ;
laciniis calycinis oblongis , erectis.
Veronica stems erect, pubescent leaves oblong, cerulate;
flowers axillary, solitary, short pedunculate, shorter than leaves;
calyx lobes oblong, erect.
Crescit in Regno Mexicano prope Xalapa (alt. 630 hex.), in nemoribus Liquidambaris
Styracifluae. ☉ Floret Febrruario.
He grows in the Mexican kingdom near Xalapa (alt. 630 fathoms [=3780 ft.]), in the woods of the Liquidambar styracifluae [American sweetgum]. Annual. It blooms in February.
Longer Latin description omitted.
Differt a Veronica peregrina ; cui simillima, caule pubescente, foliis latioribus ,
floribus pedunculatis laciniisque calycinis latioribus.
It differs from Veronica peregrina; which it most resembles, the stem is pubescent,
the leaves wider, flowers pedunculate, with broad calyx lobes.
- Pennell, Francis W., 1919c.
7a. Veronica peregrina xalapensis (H. B. K.) Pennell, comb. nov.
Veronica xalapensis H. B. K., Nov. Gen. et Sp. 2: 389.
1817. “Crescit in Regno Mexicano prope Xalapa
(alt. 630 hex.), in nemoribus Liquidambaris Styracifluae.”
Occasional in cultivated soil. In the western half of the continent
this glandular-pubescent plant completely replaces true
peregrina. In the east it is only occasionally seen, and that
probably as an introduction. Intergradation to the species
seems to be complete.
• Golden Checklist Flora:
Galium aparine;
Galium aparine L. “Sticky Willy, Cleavers”
• Golden Checklist Flora:
Galium boreale;
Galium boreale S. Watson “Bedstraw”
• Golden Checklist Flora:
Galium triflorum;
Galium triflorum Michx. “Fragrant Bedstraw”
• Golden Checklist Flora:
Non-native Caprifoliaceae;
Caprifoliaceae in the Golden Flora
• Golden Checklist Flora:
Dipsacus fullonum;
Dipsacus fullonum L. “Fuller's Teasel”

“Fuller's Teasel” at Tin Cup Ridge, Coll. No. 2677.
Coll. No. 2677, Dipsacus fullonum
- Linne´, Carl von, 1753.
Original Text
1. DIPSACUS foliis feffilibus ferratis.
fullonum.
Dipfacus foliis connato-perfoliatis,
Hort. upf. 25.
ariftis fructus rectis.
Sauv. monfp. 156.
Dipfacus capitulis florum conicis.
Hort. cliff. 29.
Gron. virg. 15.
Roy. lugdb. 188.
Dalib. parif. 44.
Dipfacus fylveftris aut Virga Paftoris major.
Bauh. pin. 385.
Dipfacus fylveftris.
Dod. pempt. 735.
β Dipfacus fativus.
Bauh. pin. 385.
ariftis fructus hamatis.
Sauv. monfp. 156.
Habitat in Gallia, Anglia, Italia.
♂
• Golden Checklist Flora:
Dipsacus laciniatus;
Dipsacus laciniatus L. “Cutleaf Teasel”
- Linne´, Carl von, 1753.
Original Text
2. DIPSACUS foliis connatis finuatis.
laciniatus.
Dipfacus folio laciniato.
Hauh. pin. 385.
Morif. hift. 3. p. 158. f. 7, t. 36. f. 4,
Habitat in Alfatia, Azow.
♂
• Golden Checklist Flora:
Knautia macedonica;
Knautia macedonica Griesb. “Macedonian scabious”
- Griesbach, August, 1839.
Frequens in omni Macedonia occidentali alt. 800'-3000',
ubi in fruticetis consociata crescit cum Kn. hybrida: in campo
Bitoliensi, pr. Perlepe, Ueskueb, copiose in campo Tettovo
(substr. var.)! Fl. Jul. Aug. E.
Frequent in all western Macedonia alt. 800'-3000',
where it grows in shrubs associated with Kn. hybrida: in the field
of Bitola, at Perlepe, Ueskueb, copiously in the field of Tettovo
(substr. var.)! Fl. Jul. Aug. E.
• Notes on Colorado Flora:
Lonicera involucrata;
Lonicera involucrata (Richardson) Banks ex Spreng. “Twinberry Honeysuckle”
• Golden Checklist Flora:
Lonicera morrowii;
Lonicera morrowii A. Gray “Morrow's honeysuckle”
- Gray, Asa, 1856.
Original Text
This collection, hastily made as their small opportunities permitted, by Dr. Morrow and my
early friend, Mr. Williams, of the American Mission at Macao, has proved a very interesting
one, as might have been expected. No part of the world beyond his own country offers, as to
its vegetation, a greater interest to the botanist of the United States than Japan. This is not
the place to dwell upon the very remarkable relations which subsist between the Flora of Japan
and that of the United States, which have been more or less noticed ever since Thunberg's Flora
Japonica was published. As the present collection was principally made in the northern part of
Japan, it was naturally anticipated that it would add to the Japanese Flora a considerable
number of species identical with, or closely allied to, those already known to inhabit Kamtschatka
and Northwest America. This proves to be the case ; and at the same time it has brought to
view an equal number of Eastern United States forms, no insignificant portion of which are
specifically identical. These interesting facts will arrest the botanist's attention as he turns
over these pages. I refrain from commenting upon these relations until another and perhaps a
larger collection of Japanese plants shall be made known, namely, that made by that excellent
and most assiduous collector, Mr. Charles Wright, in the North Pacific Exploring Expedition,
under Commodore Rodgers.
The present collection makes known nearly forty new species and one new genus ; a very
considerable number under the circumstances of the case. Thirteen of these are Carices ; but
as most of these were gathered too early in the season, it is probable that the number will be
somewhat reduced when they are better known. I have to thank my excellent friend, Dr.
Boott, of London, the most learned Caricologist of the age, for having studied the specimens of
this genus, and furnished the descriptions for this report. I have also to thank my promising
young friend, Mr. Daniel C. Eaton, for having studied the Ferns of the collection, a department
of botany to which he devotes special attention. Mr. Sullivant has obligingly determined the
Mosses, and Prof. Harvey, of Dublin, the Algas of the collection.
ASA GRAY.
Original Text
Lonicera (Xylosteum) Morrowi (n. sp.) ; erecta, molliter tomentosa; foliis oblongis basi rotundata
vel subcordata supra pubescentibus subtus canescenti-tomentosis ; pedunculis folio dimidio
brevioribus flores longitudine asquantibus ; bracteis lanceolatis vel spathulatis ovariis plerumque
discretis duplo quadruplove longioribus ; corolla ultra medium fissa basi hinc gibbosa ; filamentis
basi styloque hirsutis.
—
Hakodadi, on hill-sides.
Whole plant soft-downy, except the upper
surface of the leaves. These are about an inch and a half long, by three-fourths of an inch in
width, and on petioles a line and a half long. Corolla whitish ? somewhat pubescent ; the
lobes narrowly oblong. Bracts variable in size, sometimes five lines long, foliaceous. The
pubescence, and especially the long peduncles, must distinguish this from L. brachypoda, DC.
The specific name is a deserved compliment to one of the collectors.
• Golden Checklist Flora:
Lonicera tatarica;
Lonicera tatarica L. “Tatarian Honeysuckle”
- Linne´, Carl von, 1753.
- Stearn, William T., 2004.
Original Text
5. LONICERA pedunculis bifloris, bassis diftinctis, foliis cordatis obtufis.
Hort. npf. 42.
tatarica.
Chamaecerafus fructu gemino rubro, foliis glabris cordatis.
Amm. ruth. 184.
Habitat in Tataria.
♄
Symphoricarpos
- Bell, Charles D., 2010.
Authors for Symphoricarpos in FNANM are: Applequist, Wendy L./wendy.applequist at mobot.org and Bell, Charles D./valerianaceae1969 at gmail.com. Caprifoliaceae will be contained in Volume 18, which, as of this date, 3 August 2014, is under production.
- Bell, Charles D., 2010.
“Towards a Species Level Phylogeny of Symphoricarpos (Caprifoliaceae), Based on Nuclear and Chloroplast DNA”
• Golden Checklist Flora:
Symphoricarpos occidentalis;
Symphoricarpos occidentalis Hook. “Western Snowberry”
- Hooker, Sir William Jackson, 1829-1840.
Original Text
2. S. occidentalis ; spicis densis terminalibus axillaribusque nutantibus,
corolla intus laciniisque dense barbatis,
stylo staminibusque subexsertis.
—
Rich. in Frankl. 1st Journ. ed. 2. App. p. 6.
Hab. Woody country, between lat. 54° and 64°,
and known under tbe name of “Wolf-berry.” Dr. Richardson.
Abundant about the Saskatchawan and Red River. Drummond. Douglas.
Fort Vancouver, on tbe Columbia. Douglas.
—
Dr. Richardson has justly remarked of this plant “S. racemosi proximus:”
and among the numerous specimens in the Herbarium,
are some which appear almost as much allied to the
one species as tbe other.
But the majority of individuals of the two species are readily enough distinguished ;
those belonging to the S. occidentalis, by their larger, less glaucous, more rigid, and denser foliage,
(some of tbe leaves being two and a half inches long,)
by the flowers arranged in dense drooping spikes, larger than in S. racemosus,
and by the prominent style and stamens.
• Golden Checklist Flora:
Symphoricarpos rotundifolius;
Symphoricarpos rotundifolius A. Gray “Roundleaf Snowberry”
- Gray, Asa, 1852-1853.
Symphoricarpos rotundifolius (sp. nov.) : foliis orbiculatis seu ovato-rotundis
parvulis ramulisque molliter pubescentibus ; floribus axillaribus solitariis ; bracteolis
ovario brevioribus dentibusque calycis cyathiformis ovatis obtusis glabris ; corolla
infundibuliformi intus glabra, lobis stamina superantibus.
—
Sides of mountains
around the copper mines, New Mexico ; Aug. (1388.)
—
Stems 3 or 4 feet high,
much branched. Leaves 5 to 9 lines long and 4 to 9 wide, softly and finely pubescent
above, and finely tomentose underneath, entire, or the larger often repand ; the
petiole a line or less in length. Flowers almost sessile in the axils of the upper
leaves. Calyx with the ovary a line and a half long, its lobes slightly if at all
ciliate. Corolla 3 lines long, tubular-funnelform. Fruit not seen.
—
The leaves
are rounded and more downy than in Nuttall's S. mollis, and the inflorescence as
well as the shape of the corolla is entirely different. The flowers are more like
those of S. glaucescens and S. montanus, but only half as large ; and they, as well
as the leaves, are larger than in S. microphyllus (as described by Kunth), with
which in other particulars its characters do not accord.
• Golden Checklist Flora:
Viburnum lantana;
Viburnum lantana L. “Wayfaringtree”
- Linne´, Carl von, 1753.
Original Text
Lantana.
5. VIBURNUM foliis ovatis ferratis venofis fubtus tomentofis
Vir. cliff. 25.
Hort. upf. 68
Roy. lugdb. 242.
Sauv. monfp. 136.
Viburnum. Hort. cliff. 107.
Viburnum vulgo. Bauh. pin. 249.
Lantana. Dod. pempt. 701.
Habitat in Europae auftralioris fepibus afgillosis.
[He lives on the hedges of the south of Europe.]
♄
• Golden Checklist Flora:
Sambucus canadensis;
Sambucus canadensis L. “American Black Elderberry”
- Linne´, Carl von, 1753.
• Golden Checklist Flora:
Campanula glomerata;
Campanula glomerata L. “Dane's Blood”

Coll. No. 1982, Campanula glomerata
- Linne´, Carl von, 1753.
Original Text
glomerata.
19. CAMPANULA caule angulato finplici floribus
feffilibus, capitulo terminali.
Vir. cliff. 16.
Fl.fuec. 188.
it. gotl. 196.
it. fcan. 306.
Roy. lugdb. 245.
Dalib. parif. 67.
Campanula foliis lanceolato-ovatis crenatis, ramis
capitulo florali terminatis.
Hort. cliff. 64.
Campanula pratenfis, flore conglomerato.
Bauh. pin. 94.
Trachelium alpinum, floribus conglomeratis, foliis
afarinae rigidis & hirfutis.
Herm. par. 235. t. 235.
Habitat in Angliae, Galliae, Sueciae pratis aridis.
♃
Folia ovato-oblinga, seffilia, obtufiufcula.
Flores 3, fesfiles in alis foliorum;
captiulum terminale.
Campanula petiolata A. DC. “Western Harebell”
- Candolle, Alphonse de, 1830.
66 . CAMPANULA PETIOLATA.
66. Campanula petiolata.
C. glaberrima simplex pauciflora , foliis radicalibus longé petiolatis cordatis rotundatis dentatis ,
petiolis marginatis , caulinis inferioribus lanceolatis remotč dentatis petiolatis , superioribus linearibus
integris , lobis calycinis erectis subulatis corollâ campanulatâ duplň brevioribus.
C. glaberrima simplex pauciflora,
radical leaves with long petioles, cordate, round, toothed, marginate petioles,
lower stems, lanceolate, remotely toothed, upper ones, linear, entire,
erect calyx lobes subtended by corolla, campanulate, double shorter.
Habitat in Americâ boreali propč lacum dictum Slave-Lake. (Vid. in herb. Hook. )
He lives in North America near the lake called Slave-Lake.
(I have seen it in the herbarium of Hooker.)
Planta habitus C. rotundifolić. Radix tenuis. Caules pedales , erecti , simplices , cylindrici ,
teretes , ut tota planta glaberrimi. Folia radicalia parům numerosa, magna, longč petiolata ,
petiolis 1-2 poll. longis , sćpč marginatis , supernč latis , in limbos ovato-rotundatos , subcordatos ,
remotč dentatos vel subintegros , semipollicem latos , expansis : caulina inferiora ovata et lanceolata ,
lata , serrata , petiolata , supernč magis ac magis angustiora , linearia et integra , ut in vulgari
C. rotundifoliá. Flores 3-5 circiter, apice plantć , pedicellati , pedicellis unifloris, erectis
post anthesin incurvatis. Calyx 5-fidus , glaber, tubo obconico ; lobis subulatis, erectis, 3 lin.
longis. Corolla cćrulea ; campanulata, lobis calycinis duplo longior. Stylus ut in C. rotundifoliá.
Capsula nutans , ovoidea , 3 lin. longa , lobis calycinis erectis terminata.
Plant in the habit of C. rotundifoliae.
The root is thin. Stems pedal, erect, simple, cylindrical, smooth, like the whole plant very glabrous.
Radical leaves few in number, large, long petiolate, petioles 1-2 inch. long, often emarginate, uppermost broad, in ovate-rounded limbs, subcordate,
remotely toothed or subentire, half an inch wide, spreading:
lower stem ovate and lanceolate, broad, serrated, petiolate, uppermost more and more narrow, linear and entire, as in vulgaris C. rotundifolia.
Flowers about 3-5, at the apex of the plant, pedicellate, single-flowered pedicels, erect and curved after anthesis.
Calyx 5-fidus, glabrous, obconic tube; lobes subulates, erect, 3 lin. long
Corolla blue; bell-shaped, twice as long as the lobes of the calyx.
Style as in C. rotundifoliá.
Capsule nodding, ovoid, 3 lin. long, terminated by erect lobes of the calyx.
Doct. Richardson in peregrinatione trans Americani borealem propč laçum Servi (Slave-Lake)
detexit. Affinis est C. rotundifolić , sed ab eâ differt , foliis radicalibus majoribus limbo minůs
dentato-crenato , petiolo supernč marginato. A C. linifoliâ iisdem characteribus et floribus minoribus ,
foliisque radicalibus magěs cordatis. Parům distat ŕ varietate Langsdorffianâ , illius
speciei, in eisdem regionibus habitante. Florem similem habet, et caulem ut in eâ rectum et
pauciflorum ; sed planta hic descripta foliis radicalibus majoribus , dentatis , cordatis , petiolis
marginatis, ab eâ varietate vel specie tamen aliquatenůs differt.
Dr. Richardson, in his travels across America, discovered the boreal lake near Slave-Lake.
It is related to C. rotundifoliae, but it differs from it, with larger radical leaves,
a smaller toothed-crenate margin, and a marginate upper petiole.
A C. linifolia with the same characters and smaller flowers, and more cordate radical leaves.
It is a little distant from the Langsdorffian variety, of that species, inhabiting the same regions.
It has a similar flower, and a stem that is straight and few-flowered;
but the plant here described, with larger radical leaves, toothed, cordate, and margined petioles, differs somewhat from that variety or species.
• Golden Checklist Flora:
Campanula rapunculoides;
Campanula rapunculoides L. “Rampion Bellflower”
- Linne´, Carl von, 1753.
Original Text
rapunculoides.
14. CAMPANULA foliis cordato-lanceolatis, cauleramofo,
floribus fecundis fparfis.
Vir. cliff. 17.
Roy. lugdb. 246.
Dalib. parif. 66.
Campanula foliis ovatis crenatis, floribus per caulem
uno verfu fparfis.
Hort. cliff. 64.
Campanula hortenfis, rapunculi radice.
Bauh. pin. 94.
Morif. hift. 2. p.f. 5. t. 3. f. 32.
Habitat in Helvetia, Gallia.
• Golden Checklist Flora:
Campanula rotundifolia;
Campanula rotundifolia L. “Harebell”
Original Text
3. CAMPANULA foliis radicalibus reniformibus,
caulinis linearibus.
Fl. lapp. 83.
Fl. fuec. 176
Hort. upf. 65.
Roy. lugdb. 247.
Dalib. parif. 66.
rotundifolia.
Campanula minor rotundifolia vulgaris.
Bauh. pin. 93.
β Campanula minor rotundifolia alpina.
Bauh. pin. 93.
prodr. 34. t. 34.
γ Campanula alpina linifolia caerulea.
Bauh. pin. 93.
Fl. lapp. 84.
it. gotl. 229.
Habitat in Europae pafcuis.
♃
- Sutherland, Brittany L., and Laura F. Galloway, 2018.
- Herrando-Moraira, Sonia, et al., 2019.
Tribe Cardueae
• Golden Checklist Flora:
Achillea millefolium;
Achillea millefolium L. “Common Yarrow”
- Linne´, Carl von, 1753.
Original Text
Interpretation and Comments
14. ACHILLEA foliis bipinnatis nudis:
laciniis linearibus dentatis.
Hort. cliff. 413.
Fl. fuec. 705.
Mat. med. 397.
Roy. lugdb. 175.
Gron. virg. 101.
Dalib. parif. 263.
Achillea foliis pinnato-pinnatis. Fl. lapp. 311.
Millefolium vulgare album. Banh. pin. 140.
β. Millefolium purpureum. Tabern hift. 130.
Millefolium.
Habitat in European meadows and pastures.
- Kalm, Pehr, 1762.

Kalm (1762) mention of Achillea millefolium in Montreal, Canada.
- Pursh, Frederick, 1814.
Original Text
Interpretation and Comments
3. A. foliis bipinnatifidis pilosis linearibus dentatis mucronatis,
caulibus sulcatis.
Smith fl. brit. 2. p. 908.
Willd. sp. pl. 3. p. 2208.
Millefolium.
Probably need to see what Willdenow says about A. millefolium, i.e., does he mention North American occurrence?
However, when I checked (11 May 2021),
the Real Jardin Botanico (Madrid) server was not working.
- Nuttall, Thomas, 1818.
- Torrey, John, and Asa Gray, 1838-1843.
Agoseris sp., “Goat-Chicory”
- Schreber, Johann Christian Daniel van, 1791.
• Publication Details:
Schreber, 1789-1791, publication details;
- Robin, C. C. (Charles-Ce´sar), 1817.
Rafinesque gets/takes credit for the name Agoseris.
Original Text
English Translation
Comments
180. Troximon odoratum Raf.
Hispidum, ramis fastigiatis, foliis sessilibus ovato-oblongis acutis integris ciliatis scabris,
floribus racemosis nudis, perianthis pilosis 8 partitis, sub 12 floris Raf.
—
Chicoracee fenouillette Rob. P. 425.
Stems four feet high, round milky;
leaves a little thick,
flowers pale yellow,
sweet-scented,
ligules five-toothed, anthers and stigmas very projecting, deep yellow ;
seeds oblong compressed striated with a thick long down.
This species together with Tr. Virginicum, Tr. Pallidum and Tr. Bulbosum
will form the genus Troximon ;
the other species which are acaules and with an embricated calyx,
must form a peculiar genus which I shall call Agoseris. Raf.
Found in woods.
The way I read this, Rafinesque has validly published Agoseris
by saying that it is like Troximon that is
acaulescent and has an imbricated calyx (or the involucral bracts are imbricated).
- Nuttall, Thomas, 1840-1841.
• Glossary:
rostrum;
• Taxa Notes:
Troximon glaucum, Nuttall, 1841;
Agoseris parviflora, Nuttall, 1841;
Original Text
Comments
TROXIMON. (Nutt. Gen. Am., non Gaertner)
Capitulum many-flowered.
Involucrum imbricate, subcampanulate, divisions lanceolate, distinct, or united at base.
Receptacle naked, punctate.
Achenium subterete, with ten obtuse ribs, attentuated above into a somewhat similarly
striated, and rather short, thick rostrum.
Pappus copious, setaceous, persistent, widest at base, longer than the achenium,
and scarcely scabrous.
—
Stemless perennials, with fusiform roots, and mostly entire, linear, smooth,
sublanceolate leaves.
Scapes terete, exserted, one-flowered; flowers yellow or rose-coloured.
OBS. The only species of this genus known to Gaertner, T. lanatum,
is now referred to Scorzonera,
the name thus unoccupied may, therefore, still be retained for the American species.
The question is: was Troximon really available?
Or, was Troximon illegitimate because Krigia had priority?
Nuttall then goes on to propose Troximon glaucum (=Agoseris glauca)
and T. parviflorum sp. Nov. (=A. parviflorum).
- Lee, Joongku, Bruce G. Baldwin, and L. D. Gottlieb, 2003.
• Golden Checklist Flora:
Agoseris glauca;
Agoseris glauca (Pursh) Raf. “Pale Goat-Chicory”
- Moulton, Gary E., 1999.
- Pursh, Frederick, 1816.
• Publication Details:
Pursh, 1814, publication details;
Original Text
Translation and Interpretation
Comments
604. TROXIMON. Gaert. Carp. P. 360. Pers. Syn. 2 p. 360
1. T. scapo unifloro, calycinis foliolis imbricatis cuspidatis,
foliis linearibus integerrimis utrinque glaucis.
glaucum.
Troximon with single-flowered scape, involucre with imbricate bracts with acute tips, green leaves are linear with entire margins.
glaucum.
On the banks of the Missouri.
♂.
v. s. ;
v. v. in Hortis.
Flowers bright yellow.
On the banks of the Missouri.
Biennial.
I have seen it dried.
I have seen it live in gardens.
Flowers bright Yellow.
Pursh doesn't tell us whose dried collection he saw.
It could have been a Bradley collection.
It probably was not a Lewis & Clark collection because Moulton (1999) does not list an Agoseris collection in the
Lewis & Clark herbarium.
• Glossary:
rostrum;
• Taxa Notes:
Troximon, Nuttall, 1841;
• Publication Details:
Nuttall, 1841, publication details;
Original Text
Comments
†
Achenium terete, shortly rostrate, with obtuse ribs
This is the first section of Nuttall's Troximon and will contain T. glaucum [=Agoseris glauca].
Troximon glaucum.
The involucrum is usually smooth,
the divisions in about three series,
the outer shorter, all of them lanceolate and acute.
HAB. On the plains of the Platte, and Missouri, about the Great Bend.
- Dietrich, David Nathaniel Friedrich, 1847.
• Publication Details:
Dietrich, 1847, publication details;
3606. AGOSERIS Rafin. (1817).
(Troximon Nutt.)
2. A. cuspidata; …
2. A. glauca;
glaucescens;
fol. lineari-lanceolatis acutus integris scapoque glabriusculis;
setis pappi capillaribus rigidis.
Troximon Nutt. B. M. T, 1667.
β dasycephalum Torr. Et Gray.
T. glaucum α Hook. B. M. t 3462.
Ammogeton scorzonerifolium Schrad. In Amer. Bor.
♃.
3 A. parviflora (Troximon Nutt.): …
4. A. rosea …
5. A. taraxacifolia; …
• Golden Checklist Flora:
Agoseris parviflora;
Agoseris parviflora (Nutt.) D. Dietr. “Steppe Goat-Chicory”

Coll. No. 1101, Agoseris parviflora
- Nuttall, Thomas, 1840-1841.
• Taxa Notes:
Troximon, Nuttall, 1841;
• Publication Details:
Nuttall, 1841, publication details;
Original Text
Comments
†† Achenium compressed, with ten shallow, acute ribs, and attenuated into a distinct rostrum,
shorter than the long and bristly pappus.
Involucrum ovate, in about three series of unequal, lanceolate sepals.
This is the second section of Nuttall's Troximon and it has a “distinct rostrum” or beak.
The section starts off with Nuttall's new species T. parviflorum [=Agoseris parviflora].
Troximon * parviflorum;
leaves linear-lanceolate, acuminate, smooth or pubescent,
often runcinately denticulate towards the base;
scape pubescent at the summit, lanuginous;
sepals nearly smooth, in three unequal series, lanceolate, acuminate;
flowers yellow.
Hab. On the plains of the Platte to the Rocky Mountains.
About four or five inches high.
The leaves about two lines wide, acuminated at each end.
Pappus minutely scabrous;
the outermost divisions of the involucrum only about half the length of the inner.
- Dietrich, David Nathaniel Friedrich, 1847.
• Publication Details:
Dietrich, 1847, publication details;
3606. AGOSERIS Rafin. (1817).
(Troximon Nutt.)
. A. cuspidata; …
2. A. glauca; …
3 A. parviflora (Troximon Nutt.):
glabriuscula;
fol. Anguste lanceolato-linearibus acute asuminatis integris subretrorsp-denticulatis;
pappo capillari rigido.
In mont. Rocky.
♃.
4. A. rosea …
5. A. taraxacifolia; …
• Golden Checklist Flora:
Ambrosia acanthicarpa;
Ambrosia acanthicarpa Hook. “Flat-Spine Burr-Ragweed”
• Golden Checklist Flora:
Ambrosia artemisiifolia;
Ambrosia artemisiifolia L. “Annual Ragweed”
• Golden Checklist Flora:
Ambrosia psilostachya;
Ambrosia psilostachya DC. “Western Ragweed”
- DeCandolle, Augustus Pyramus, 1836.
Original Text
9. A. psilostachya,
caule herbaceo erecto tereti paniculato-ramoso foliisque
scabridis, foliis subsessilibus pinnatipartitis lobis lineari-lanceolatis acnminatis
hinc inde pinnatim incisis, racemis plurimis gracilibus inter capitula mascula
remota ebracteatis, fructibus infra apicem tuberculis obtusis 5-6 onustis apice
conico elongato.
—
in Mexico inter San-Fernarido et Matamoros legit cl.
Berlandier (pl. exs. n. 2280!).
Fructus plurimi parvi in invol. foem. Capitula
masc. pubero-scabrida parva. (v. s.)
• Golden Checklist Flora:
Ambrosia tomentosa;
Ambrosia tomentosa Nutt. “Ragweed”
- Nuttall, Thomas, 1818.
• Publication Details:
Nuttall, 1818, publication details;
603. AMBROSIA. L. (Bitter-weed.)
Monoicous (sic).
—
Masc.
Calix 1-leaved.
Anthers approximate, but not united.
Receptacle naked.
—
Fem.
Calix 1-leaved,
entire or 5-toothed,
1-flowered.
Corolla none.
Nut formed from the indurated calix, 1-seeded.
Tall herbaceous and mostly annual plants;
leaves rough, the lower most opposite, the upper alternate,
bipinnatifid, trifid, or rarely entire;
flowers in long terminal and proximately axillar spikes,
upper flowers masculine numerous,
the lower fewer, feminine, glomerated, clusters 2 to 5-flowered, tribracteate.
Species.
1. A. integrifolia.
2. bidentata.
3. trifida.
4. elatior.
5. artemisifolia.
6. paniculata.
7. heterophylla.
8. * tomentosa.
Perennial; stem low;
leaves bipinnatifid, upper side white and tomentose;
spikes solitary.
Hab.
In Upper Louisiana on the banks of the Missouri; rare.
Only 1 or 2 feet high.
A North American genus, with the exception of 1 species in Peru
and another indigenous to the sea-coasts of the Levant.
- Nuttall, Thomas, 1840-1841.
• Publication Details:
Nuttall, 1841, publication details;
FRANSERIA. (Cavan.)
Franseria * bipinnatifida; ...
Franseria * pumila; ...
Franseria * discolor;
♃, root creeping;
leaves ubterruptedly bipinnatifid,
above nearly smooth, canescently and closely tomentose,
segments subovate, acute, confluent in the wide rachis;
stem short, with the lateral branched decumbent.
Hab.
In the Rocky Mountains, neat the Colorado of the West.
A very remarkable and distinct, as well as elegant species.
Stem about a span long, slightly pubescent;
leaves on long petioles, with a lanceolate outline, acute,
about six inches lone, white beneath, green above,
the pinnatifid segments lanceolate, the rachis incisely toothed.
Male florets rather numerous;
receptacle with narrow, pubescent palea;
involucrum about five or six-toothed;
female flowers few, fruit spiny.
Franseria * cuneifolia; ...
• Golden Checklist Flora:
Ambrosia trifida;
Ambrosia trifida L. “Giant Ragweed”
• Golden Checklist Flora:
Anaphalis margaritacea;
Anaphalis margaritacea (L.) Bentham & Hooker “Western Pearly Everlasting”
Original Text
margaritace-
um.
3. GNAPHALIUM foliis lineari-lanceolatis acuminatis
alternis, caule fuperne ramofo, corymbis faftigiatis.
Hort. cliff. 401.
Hort. upf. 255.
Gmel. fib. 2. p. 107.
- DeCandolle, Augustus Pyramus, 1836.
- Weber, William A., and Ronald C. Wittmann, 2012.
• Golden Checklist Flora:
Antennaria parvifolia;
Antennaria parvifolia Nutt. “Small-Leaf Pussytoes”

Habitat of Coll. No. 1128, Antennaria parvifolia
Coll. No. 1909, Antennaria parvifolia
- Nuttall, Thomas, 1840-1841.
• Glossary:
sarment;
• Publication Details:
Nuttall, 1841, publication details;
Original Text
Antennaria * parvifolia;
subcaespitose, with procumbent sarments;
stem simple;
lower leaves spathulate, or spathulate-linear, the upper linear, all whitely tomentose;
flowers conglomerate;
scales of the involucrum oblong-ovate, eroded, yellow.
Hab.
On the Black Hills and plains of the upper part of the Platte.
A dwarf species, spreading out in canescent tufts with very small leaves,
which are about half or three quarters of an inch long, and about two or three lines wide;
the flowers in an irregular, somewhat round mass, not a circular corymb,
with the scales of the corynb sulphur yellow, and very conspicuous.
Radical leaves somewhat rhomboidally spathulate.
The pappus of the male flower is very conspicuously clavellate:
the female flower has purple oblong-lanceolate scales to the involucrum,
and a filiform pappus.
A specimen of this sex from Altai has a near resemblance to our plant,
but is larger in all its parts,
and is the A. hyperborea of Don.
- Bayer, Randall J., 1990.
• Golden Checklist Flora:
Antennaria rosea;
Antennaria rosea Greene “Rosy Pussytoes”
- Greene, Edward L., 1898.
Original Text
A. rosea. A. dioica, var. rosea, Eaton, Bot. King Exp. 186.
A. parvifolia, var. rosea, Greene, p. 175 supra.
A. parvifolia, Nutt. in small part, but not of spec. char. Plant by
no means small, often 12, sometimes 14 or 15 inches high,
yet frequently only 6 or 8 inches: leaves comparatively
small, of the thinnest, as to texture, only canescently tomentose,
but permanently so on both faces, the quite gradually
dilated upper portion acute; cauline long and narrow, acute
or acuminate: heads small, closely compacted
pound cymose rounded cluster: bracts of the involucre
pluriserial, their basal part concealed by wool, the tips from
broad and obtuse in the outer series to narrow and acute
in the inner, all rose-red.
Of this only the female plant is known to me; which is
the more remarkable in view of the fact that no other
north-western Antennaria so abounds in every large herbarium.
It is a dry-ground species of subalpine habitat, with either
short or elongated slender dry and subligneous stolons; in
this quite unlike the true A. parvifolia. From A. hyperborea
more difficult to distinguish it, except by the looser
inflorescence, longer heads, less woolly involucre, and fewer
paler narrower bracts of the last named. I give the following
rather copious list of localities for A. rosea.
North Park, Colorado, Chas. S. Sheldon, n. 128 ;
Yellowstone Park, Frank Tweedy, n. 728;
mountain meadows in Kootenai Co., Idaho, J. B. Leiberg, n. 646 ;
Nez Perces Co., Idaho, Heller, n. 3441 ;
Salmon River, British Columbia, Dawson (Can. Surv. n. 11281);
Spence's Bridge, B. C, Macoun, n. 11282;
summit of Mt. Arrowsmith, Vancouver Island, Macoun, n. 11279;
pine woods near Spokane, Washington, C. V. Piper, n. 2273;
Mt. Hood, Oregon, Thos. Howell;
Crooked Creek, southeastern Oregon, and
Warner Range, northeastern California, Mrs. Austin.
• Golden Checklist Flora:
Arnica cordifolia;
Arnica cordifolia Hook. “Heart-Leaf Leopardbane”
- Hooker, Sir William Jackson, 1829-1840.
Original Text
Comments and Interpretation
4. A. cordifolia ; caule elato,
foliis cordatis glabriusculis basi sinu distincto,
inferioribus longe petiolatis pari supremo solummodo sessili,
panicula 1-3-flora.
Hab. Alpine woods of the Rocky Mountains, on the east side, Drummond ;
and on the west side,
in mountain woods between the Kettle Falls and Spokan River,
and in the Blue Mountains. Douglas.
—
This appears quite distinct from the preceding,
and is very constant in the shape and petiolation of its leaves.
The preceding was A. menziesii.
• Golden Checklist Flora:
Arnica fulgens;
Arnica fulgens Pursh “Shining Leopardbane”
- Pursh, Frederick, 1814.
Original Text
Comments
3. A. pubescens ; foliis radicalibus lanceolatis obtusiusculis
basi attenuatis petiolatis trinervibus, caulinis oppositis
remotis linearibus, caule unifloro.
fulgens.
Pursh has seen a dried specimen, but does not say whose specimen or where he saw it.
- Nuttall, Thomas, 1818.
Original Text
563. ARNICA. L.
Calix hemispherical, leaflets equal, mostly in
a simple series? Radial florets often producing
5 filaments destitute of anthers. Receptacle naked.
Pappus simple, scabrous.
A polymorphous and divided genus? Some of the species
caulescent and also shrubby; those of Europe and
North America, with a few others, often scapigerous,
scapes 1-flowered, sometimes producing 1 or 2 pair of
opposite leaves; flowers mostly yellow.
Species. 1. A. montana. β fulgens.
A. fulgens. Ph. 2. p. 527.
Scarcely dissimilar to specimens of the alpine
variety in the Banksian herbarium.
Hab. On the margins
of marshy springs and in depressed situations, from the
Arikarees to Fort Mandan, and probably as far as the
Mountains. Flowering in July. Flowers bright yellow.
Obs. Minutely pubescent. Scape about 12 inches high,
mostly with 2 pair of leaves, terminated by 1 rarely 3
flowers. Leaves oblong-lanceolate, 3 to 5 nerved; summit
of the caudex tomentose. Calix a simple series of
leaflets, somewhat hirsute. Tube of the florets pilose;
rays without filaments; seed hirsute, slender.
A. plantaginea, of Pursh from the specimen
which I have seen, appears also a mere variety of the above.
2. Doronicum. 3. nudicaule. Doroonicum nudicaule. Mich.
2. p. 121. A. Claytoni. PH. 4. maritima.
A genus principally indigenous to Europe and the Cape
of Good Hope, there are also 2 species in South America,
2 in Japan, 1 in Arabia Felix, and a shrubby species, or
something else, in New Zealand.
- Riddell, John L., 1835.
848. Arnica fulgens, Pursh.
Variety of A. montana, Nutt. Leopard's bane.
Jul. y. 1 f.
♃.
Margins of Marshy springs. Mo. Ter. Nutt.
Stimulant, diaphoretic, emmenagogue, &c.
Wood and Bache. 108.
Artemisia L.
Artemisia Linnaeus subg. Absinthium DC.
One taxon in Artemisia subgroup Absinthium is not recognized by Ackerfield (2015)
- Ackerfield, Jennifer, 2022.
Artemisia subg. Artemisia L
Artemisia Linnaeus subg. Dracunculus Besser, Bull. Soc. Imp. Naturalistes Moscou. 1: 223. 1829.
- Ackerfield, Jennifer, 2015.
- Ackerfield, Jennifer, 2022.
Artemisia subg. Tridentatae (Rydberg)McArthus
• Golden Checklist Flora:
Artemisia absinthium;
Artemisia absinthium L. “Common Wormwood”
- Linne´, Carl von, 1753.
Abfinthium
12. ARTEMISIA foliis conpofitis multifidis, floribus
fubglobofis pendulis: receptaculo villofo.
Hort. cliff. 404.
Fl. fuec. 670, mat. med. 386. Roy. lugdb. 153.
Abfinthium incanum, foliis compofitis latiufcule multifidis,
floribus fubglobofis pendulis.
Gmel. fib. 3. p. 129. t. 63.
Abfinthium ponticum f. romanum, officinarum f. diofcoridis.
Bauh. pin. 138.
Abfinthium. Cam. epit. 452.
Habitat in Europae ruderatis aridis. ♃
• Golden Checklist Flora:
Artemisia campestris;
• Taxa Notes:
Notes on Artemisia campestris ssp. purshii;
Notes on Artemisia dracunculus, Linnaeus, 1753;
Artemisia campestris L. “Field Sagewort”
- Linne´, Carl von, 1753.
Original Text
Comments and Interpretation
campeftris.
4. ARTEMISIA foliis multifidis linearibus,
caulibus procumbentibus virgatis.
Hort. cliff. 403.
Fl. fuec. 668.
Roy. lugdb. 154.
Gmel. fib. 2. p. 117.
- Michaux, Andre, 1803.
CAUDATA.
A. erecta, glabra :
foliis subsetaceo-pinnatifidis ; laciniis convexis :
caule primariisque ramis in racemum longissimum , strictum dispositis ;
ramulis confertis ;
floribus omnibus pedicellatis, subgloboso-ovoideis.
A. erect, glabrous:
leaves subsetaceous-pinnatifid; lacini convex:
stem and primary branches arranged in a very long, tight raceme;
branches densely packed;
flowers all pediceled, subglobose-ovoid.
"Subsetaceous" is a term used in botanical and entomological descriptions to mean
somewhat setaceous or partially bristle-like.
 
Hab. ad ripas sabulosas fluminis Missouri.
Lives on the sandy banks of the Missouri River.
- Nuttall, Thomas, 1840-1841.
Artemisia * Pacifica; herbaceous, soft, and canescently sericeous; leaves
pseudopetiolate, bipinnately dissected, the segments often trifid, oblong or
linear, acute;
stem leaves pinnately dissected, pseudostipulate on the infertile shoots;
stem and ovate, pedicellate capitulum, smooth.
Hab.
Shores of the Pacific, at the outlet of the Oregon, in sandy places.
Considerably allied to A. canadensis, but more tomentose,
with broader and fewer segments to the leaves, the radical and
lower leaves very much as in the last species.
Perhaps A. desertorum, γ. Scouleriana of Hook.
Flor. Bor. Am., p. 325.
- Hall, Harvey M., and Frederick E. Clements, 1923.
21b.
ARTEMISIA CAMPESTRIS PACIFICA (Nuttall).
—
Root perennial; stems several or
numerous, 4 to 6 dm. high, erect from a spreading base, densely leafy below, moderately
leafy up to the inflorescence; leaves twice pinnately divided into narrowly linear divisions
about 1 mm. wide, canescent or silky pubescent, seldom if ever glabrate; inflorescence
an elongated panicle of numerous closely ascending branches, 3 to 12 em. broad or
occasionally narrowed to 1 cm.; heads subsessile, horizontal or erect, hemispheric;
involucre 2 to 3 mm. high, about 2.5 mm. broad, glabrous or sparingly short-hairy,
greenish; disk-flowers 10 to 25, the corolla 1.8 to 2.5 mm. long.
2lc.
ARTEMISIA CAMPESTRIS CAUDATA (Michaux).
—
Root biennial, often very large
when growing in sand-dunes, but perhaps never perennial; stem usually single unless
injured, erect, 3 to 5 dm. high, leafy at base and up to the inflorescence or sparsely leafy
above; leaves twice or thrice pinnately divided into elongated diverging filiform or very
narrowly linear divisions 0.5 mm. or less wide, the upper ones more simply cut, glabrous
or sometimes canescent when young (in A. forwoodi Watson, minor variation 18) ;
inflorescence a narrowly pyramidal panicle of ascending rather close branches, 3 to 10 cm. broad;
heads peduncled, nodding or erect, narrowly hemispheric; involucre 2 to 3 or rarely 3.5
mm. high, about as broad, glabrous, yellowish green; disk-flowers 5 to 15, the corolla
1.5 to 2.2 mm. long.
- Palmer, Ernest J, and Julian A. Steyermark, 1935.
Artemisia campestri L. var. caudata (Michx.) Palmer & Steyermark, comb. nov.
Wild Wormwood.
Artemisia caudata Michx. (G), (B & B).
Artemisia campestris subsp. caudata (Michx.) Hall & Clements.
Glades, bluffs, and gravel bars of streams.
Oxylophile.
Eastern Mo., south of the Missouri River: St. Louis, Jefferson, Dent, and Shannon counties.
- Peck, Morton Eaton, 1941.
Var. pacifica (Nutt.) n. comb.
Densely silky-cillous;
stems clustered in a stout caudex, spreading at base, 3-6 dm. high;
leaves bipinnately divided into linear segments about 1 mm. wide, the basal very numerous;
panicles long, narrow, and desnse, the branches short;
heads nearly globose, 2-3 mm. high;
the involucral bracts sparsely pubescent.
—
Along the Columbia R. east of the Cascades, to Alaska, S. Dak. and N. Mex.
(Subsp. pacifica H. & C.)
- Flora of North America Editorial Committee, eds., 2006.
- Weber, William A., and Ronald C. Wittmann, 2012.
• Taxa Notes:
Notes on Oligosporus, Cassini, 1817;
- Ackerfield, Jennifer, 2015.
- Ackerfield, Jennifer, 2022.
• Notes on Colorado Flora:
Artemisia campestris ssp. purshii;
• Taxa Notes:
Notes on Artemisia campestris;
Notes on Artemisia groenlandica;
Artemisia campestris ssp. purshii (Hook.) Cronquist
- Pursh, Frederick, 1814.
spithamaea.
12. A. pumila, sericeo-pubescens ;
foliis inferioribus linearibus supernae pinnatifidis,
floralibus simplicibus,
floribus racemosis brevi-pedunculatis hemisphaericis,
calycibus scariosus
In Labrador.
Colmaster.
♃
Flower large, in proportion to the size of the plant ; receptacle naked.
- Hooker, Sir William Jackson, 1829-1840.
22. A. borealis ;
22. A. borealis ;
—
α Purshii ; sericea, cinerea;
folia radicalia lineari-lanceolata integerrima, vel apice 3-5-fida:
caulina et floralia linearia:
calathidia inferiora pedunculata, superiora subsessilia ;
periclinii squamae villosae, medio fuscescentes, flosculi nudi.
Caulis adscendens 6", violaceus, superne villosissimus.
—
α Purshii ; silky, grey;
radical leaves linear-lanceolate, very entire, or 3-5-fid at the apex:
stem and flowers linear:
lower calathidia pedunculate, upper ones subsessile;
periclinian scales villous, mid-fuscescent, small flowers naked.
Stem ascending 6", violet, very villous above.
A. spithamaea. Pursh, Fl. Am. v. 2. p. 522;
A. spithamaea. Pursh, Fl. Am. v. 2. p. 522;
- Hall, Harvey M., and Frederick E. Clements, 1923.
21d.
ARTEMISIA CAMPESTRIS BOREALIS (Pallas).
—
Root perennial; stems several,
crowded on a multicipital caudex, 1 to 3 dm. high, erect from a sometimes spreading base;
densely leafy only below, sometimes moderately leafy to midway of the inflorescence;
leaves once or twice ternately divided into linear divisions, mostly 1 to 2 mm. wide, or
with a few extra lobes along the rachis and hence somewhat pinnately divided, the upper
ones simply ternate or undivided, glabrous to somewhat silky-villous and then glabrate
at least on upper surface; inflorescence raceme-like or spike-like to loosely paniculate,
1 to 5 em. broad; lower heads peduncled, the upper sessile, mostly nodding, hemispheric;
involucre 3 to nearly 4 mm. high, 3.5 to 4 mm. broad, glabrous or nearly so, yellowish
green (or brownish), the bracts with brown medial line; disk-flowers apparently 15 to 30,
the corolla 2.2 to 2.6 mm. long.
- Cronquist, Arthur, 1950.
Artemisia campestris L. var. Purshii (Hook.) Cronquist, comb. nov.
A. spithamea Pursh, Fl. Am. Sept. 522 (1814).
A. borealis Pall. var. Purshii Hook., Fl. Bor. Am. 1:326 (1833).
- Schulz, Leila M., 2006.
Synonyms:
Artemisia campestris var. purshii (Besser) Cronquist.
Artemisia campestris var. spithamaea (Pursh) M. Peck.
Artemisia campestris var. strutziae S. L. Welsh.
Artemisia purshii (Pallas) Polyakov.
Artemisia spithamaea (Hornemann) Á. Löve & D. Löve
Oligosporus borealis.
Oligosporus groenlandicus.
- Weber, William A., and Ronald C. Wittmann, 2012.
• Golden Checklist Flora:
Artemisia dracunculus L.;
Artemisia dracunculus L. “Tarragon, Dragon Wort”
- Linne´, Carl von, 1753.
• Taxa Notes:
Notes on Artemisia campestris;
Original Text
16. ARTEMISIA foliis lanceolatis glabris integerrimis.
Hort. cliff. 403.
Hort. upf. 256.
Gmel.fib. 2. p. 126. t. 59. & 60. f. 1.
Dracunculus
- Weber, William A., and Ronald C. Wittmann, 2012.
• Taxa Notes:
Notes on Oligosporus, Cassini, 1817;
- Ackerfield, Jennifer, 2015.
• Golden Checklist Flora:
Artemisia filifolia;
Artemisia filifolia Torrey “Sand Sage, Old-Man Sagebrush”
- Torrey, John G., 1828.
Original Text
210. Artemisia ludoviciana, Nutt. gen. ii. p. 143.
Arid plains of the Platte, with all the following of Nuttall :
A. Serrata, columbiensis, longifolia,
cernua, and canadensis.
Artemisia filifolia, caule herbaceo ? ramoso ?
foliis canescenti-pubescentibus ; superioribus simplicibus, filiformibus,
et subpinnatifidis, semiteretibus ; floribus conglomeratis.
- Goodman, George J., and Cheryl A. Lawson, 1995.
• Golden Checklist Flora:
Artemisia frigida;
Artemisia frigida Willd. “Prairie Sagewort”
- Stearn, William T., 2004.
- Willdenow, Carl L., 1797-1830.
Original Text
*51. ARTEMISIA frigida. W.
A. foliis incanis pinnatis , pinnis tripartitis linearibus
acutis, floralibus pinnatis tripartitisve, caule adfcendente,
floribus globofis nutantibus. W.
Kalter Beyfus. W.
Habitat in aridis frigidis Dauuriae.
♃.
Habitu praecedenti quoad Jiguram Gmelini aliquatenus
affinis fed toto coelo diverfa.
Caules adfceudentes
femipedales vel pedales ramofi, inferne glabri, fuperne
fubcanefcentes.
Folia incana, caulina pinnata, foliolis
tripartitis linearibus acutis, quadrantem pollicis
longa fesfilia; radicalia petiolata ejusdem formae et
magnitudinis , floralia triplo minora pinnata vel tantum
tripartita fesflia.
Panicula fimplex, ramis fimplicibus longis erectis.
Flores globoji breve pedunculati nutantes maguitudine Abfinthii.
Receptaculum villofum. W.
• Notes on Colorado Flora:
Artemisia groenlandica;
• Taxa Notes:
Notes on Artemisia campestris ssp. purshii;
Artemisia groenlandica Hornem.
- Friis, IB, 1996.
Artemisia groenlandica Hornem. in Icon. fl. Dan., Pl. 1585 (1818):
"Artemisia gronlandica; [diagn.]. [No auth.]."
Mat. from the Kabesund [Kaboesund] bay in Greenland, coll. Wormskjold.
Horneman (1837: 433) stated:
“A very distinctive species first found by Lt. Wormskjold in southern Greenland.”
It is assumed that Hornemann described and named the species.
A. groenlandica is now syn. of Artemisia borealis Pall. (1776);
see Bocher et al. (1978).
- Bjork, Curtis R., 2021.
…
Prior to reviewing the types of Artemisia groenlandica and A. richardsoniana,
I was prepared
to describe a new species from mountainous regions of western North America.
However, I can find
no distinctions between these widespread western North American plants
from the types of A. groenlandica or its synonyms
…
Though I have not seen specimens in the intervening regions between western North America
and Greenland, photographs posted on-line from,
for example, Rankin Inlet, Nunavut help close the geographical gap.
…
The name A. borealis var. purshii is frequently applied to this
taxon but was published by Besser as a replacement name for A. spithamea …
In western North America, A. groenlandica occurs mostly in rocky sites, especially on
limestone and serpentine, and appears to be uncommon, though widespread. …
- Nuttall, Thomas, 1818.
• Golden Checklist Flora:
Artemisia ludoviciana;
• Glossary:
connivent;
• Publication Details:
Nuttall, 1818, publication details;
Artemisia ludoviciana Nutt. “Silver Wormwood”
553. ARTEMISIA. L. (Wormwood, Southern-wood, &c.)
Calix imbricated, scales rounded, connivent.
Rays of the corolla none.
Receptacle subvillous, or nearly naked.
Pappus none.
Shrubby or herbaceous; leaves mostly multifid, flowers often racemose.
§
1. Leaves simple.
Species.
1. A * longifolia. …
2. * serrata. …
3. * columbiensis. …
4. * Gnaphaloides. …
5. * ludoviciana.
Stem simple and herbaceous;
lower leaves incise, subpinnatifid, the upper lanceolate and en- entire(sic),
on both sides pubescent, beneath tomentose;
flowers ovate, erect and sessile;
calix pubescent, panicle simple.
—
Hab.
On the banks of the Missisippi, near St. Louis;
also on the alluvial plains of the Missouri.
Obs.
Perennial.
Stem about 2 feet high.
Lower stem leaves lanceolate,
irregularly and divaricately laciniate, segments entire,
oblong-lanceolate and acute.
6. * cernua. …
7. chinensis …
§ ii. Leaves compound; stem paniculate.
8. Sontonia? Ph. 9. sericea. …
…

Coll. No. 1281, Artemisia ludoviciana
- Graustein, Jeannette E., 1967.
- Nuttall, Thomas, 1840-1841.
- Townsend, John Kirk, 1839.
Nuttall (1841, p. 400) proposed A. incompta from a collection near Hyndman Peak, Blaine County, Idaho.
Artemisia * incompta; herbaceous ; smooth, except the under surface of the
leaves, which is a little tomentose; leaves almost simply pinnatifid, trifid or
laciniate, sessile, the segments rather broad, linear and acute; flowers paniculated;
capituli subglobose, pedicellate, erect; sepals ovate and scariose; florets
numerous, smooth.
After a good breakfast, we packed our horses, and struck back
on our trail of yesterday, in order to try another valley which
we observed bearing parallel with this, at about three miles distant,
and which we conclude must of course furnish a pass
through the mountain. Although our difficulties in returning by
the same wretched route were very considerable, yet they were
somewhat diminished by the road having been partially broken,
and we were enabled also to avoid many of the sloughs and pitfalls
which had before so much incommoded us. We have
named this rugged valley, “Thornburg's pass,” after one of our
men of this name, (a tailor,) whom we have to thank for leading
us into all these troubles. Thornburg crossed this mountain two
years ago, and might therefore be expected to know something
of the route, and as he was the only man in the company who
had been here, Captain W. acted by his advice, in opposition to
his own judgment, which had suggested the other valley as affording
a more probable chance of success. As we are probably
the only white men who have ever penetrated into this most vile
and abominable region, we conclude that the name we have given
it must stand, from priority.
- Clausen, Jens, D. D. Keck, and Wm. M. Heisey, 1940.
• Publication Details:
Clausen, Keck, and Heisey;
Representatives of Artemisia ludoviciana Nutt. are shown in figure 129.
Plant 1324-3 on the left, from near Timberline station, is referable to
A. ludoviciana ssp. incompta (Nutt.) Keck comb. nov. (A. incompta Nutt. Trans. Amer. Phil. Soc. II, 7:400, 1841).
This subspecies has passed frequently under the name A. vulgaris ssp. discolor (Dougl.) H. & C.,
which is now thought referable to a more northern species, A. Michauxiana Bess.
Subspecies incompta is the subalpine and alpine form of ludoviciana,
distinguished by the deeply and twice cut divisions of the lower leaves.
Much variation and doubtless more than a single ecotype are included in this subspecies,
which occurs in the mountains of Montana and Idaho, southward to Utah, Nevada, and California.
The remaining plants in figure 129 belong to A. ludoviciana ssp. typica Keck nom.nov.
(A. ludoviciana Gen, 2: 143, 1918).
We believe this form, except in the high mountains, where it is replaced by the preceding subspecies, and in the Northwest,
where a larger-headed form intervenes, is the principal one found from the Great Basin to the Mississippi River.
Rather locally it extends westward to the eastern flanks of the Cascades and Sierras.
It is chiefly confined to the region bounded by the thirty-fifth and fifty-first parallels, but its exact boundaries fluctuate considerably.
The Great Basin form of the subspecies came from Leevining, Mono County, California;
the Rocky Mountain plants are from the east flank of Pikes Peak, Colorado.
In 1328-1 the leaves are tomentose on both sides, and this form is widely known as A. gnaphalodes Nutt.;
in 1329-1 the leaves are prominently discolored, almost glabrous above, and densely tomentose beneath,
and this matches the type of A. ludoviciana Nutt.
We consider these forms inseparable into natural subunits,
for both sorts occur together intermittently almost throughout the range of the species and obviously intermix.
In this instance these characters do not even mark ecotypical differences.

Figure 129, Artemisia ludoviciana var. incompta
- Keck, David D., 1946.

Figure 10, Leaves of Artemisia
• Notes on Colorado Flora:
Artemisia tridentata;
Artemisia tridentata Nutt. “Big Sagebrush”

Collections of subspecies of Artemisia tridentata in Colorado
- Nuttall, Thomas, 1840-1841.
Artemisia * tridentata;
♄
canescently tomentose; leaves cuneate, three-toothed at the summit,
upper ones entire and obtuse; flowers paniculate; capitulum sessile, ovate and tomentose, small;
inner sepals scariose, linear-oblong.
• Notes on Colorado Flora:
Artemisia tridentata vaseyana;
Artemisia tridentata Nutt. subsp. vaseyana (Rydb.) Beetle “Big Sagebrush”
- Rydberg, Per Axel, 1916.
112. Artemisia Vaseyana Rydberg, sp. nov.
A shrub, 5 dm. high or more; bark yellowish or brown; leaves narrowly cuneate,
3-5-dentate at the apex, 1-3 cm. long, canescent, rather conspicuously triple-veined; heads many
in a narrow panicle, with erect short branches; involucre broadly campanulate, 5 mm. high,
3-4 mm. broad; bracts about 12, in about 4 series, canescent, the outer ones broadly ovate,
less than half as long as the innermost; inner bracts broadly elliptic, obtuse, scarious with a green
midrib; flowers 5-8; corollas trumpet-shaped, 3 mm. long, sparingly glandular-granuliferous;
style barely equaling the corolla; achenes nearly 2 mm. long.
Type collected in the state of Washington in 1889, G. R. Vasey 480 (herb. N. Y. Bot. Gard.).
DISTRIBUTION: Washington, Oregon, and western Nevada.
- Beetle, Alan A., 1959.
6(a). A. tridentata subsp. vaseyana (Rydb.) Beetle, comb. nov.
Based on A. vaseyana Rydb. North American Flora 34 (3) :283. 1916.
A. tridentata subsp. vaseyana has a range geographically distinct
from that of A. tridentata subsp. tridentata. It differs mainly
in the characters emphasized by Rydberg, namely broader involucre,
more flowers per head, and broader, and more truncate or cuneate leaves.
• Golden Checklist Flora:
Balsamorhiza sagittata;
Balsamorhiza sagittata (Pursh) Nutt. “Arrow-Leaf Balsam Root”
• Tin Cup Ridge (social trail):
at Coll. 1109;
• Field Notes:
Coll. No. 1109, 14 May 2015;
How the heck did it get to Tin Cup Ridge?

Coll. No. 1109, Balsamorhiza sagittata
- Pursh, Frederick, 1816.
Original Text
649. BUPHTHALMUM. Gen. pl. 1231.
...
sagittatum.
3. B. tomentosum ; foliis radicalibus longissime petiolatis oblongis sagittatis integerrimus subtrinervibus,
caulinis oblongis in petiolum attenuatis,
caule subtrifloro,
calycinis foliolis exterioribus disco longioribus.
On dry barren hills, in the Rocky-mountains.
M. Lewis. ♃ June, July. v. s. in Herb. Lewis.
Flowers large, yellow.
The natives eat the young stems as they spring up, raw.
- Nuttall, Thomas, 1834a.
Original Text
ESPELETIA, Humb. and Bonpl.
—
BALSAMORHIZA, Hooker.
Calyx imbricatus, subsquarrosus, foliaceus.
Colollulae radii femineae, vix dentatae.
Receptaculum planum paleaceum.
Pappus nullus.
Semina compressa planiascula, subquadrangulata.
Herba perennis, pumila, grandiflora.
Helianthi facie ; radix resinosa.
This genus has also been proposed by Dr. Hooker in his Flora Boreali-Americana,
under the name of Balsamorhiza,
for a species of Heliopsis there described.
…
66. ESPELETIA sagittata.
Tomentosa, incana, foliis radicalibus longe petiolatis cordato-hastatis
integris acutis,
caulinis paucis lineari-oblongis in periolum attenuatis,
caule subtrifloro pumilo,
calicibus foliosis squarrosis, pl. 4.
BUPHTHALMUM sagittatum, Ph. 2, p. 564.
♃
With a large yellowish root like a dock.
Radical leaves about the length of the stem,
seven or eight inches,
softly and copiously tomentose, hoary.
The stem also downy, resembling a scape,
with about two or three small leaves like bracts on its upper part.
Calyx very white and softly tomentose, leafy,
the inner leaves linear-lanceolate and somewhat acute.
Rays bright yellow, very large, from sixteen to eighteen,
bidentate at the tips.
Seed flattish, elliptic, very smooth,
and wholly devoid of any vestige of pappus.
Hab.
On the borders of Flat-Head river.
Flowering in June.
The root of this plant also, when fermented a day or two in the ground,
in a hole made for the purpose,
and heated with hot stones,
is then eaten and possesses an agreeable saccharine taste.
The stems are never employed for food.
- Nuttall, Thomas, 1840-1841.
Original Text
§ II. * ARTORHIZA.
—
Leaves entire, deltoid or cordate;
involucrum very leafy at base; rays numerous.
Receptacle flat.
Balsamorhiza sagittata.
Buphthalmum sagittatum? Pursh, Vol. II., p. 564.
Espeltia sagittata; Nutt. in Journ, Acad. Nat. Sci. Philad., Vol. VII., p. 39.
Canescently tomentose;
stem low, about one to three flowered;
radical leaves cordate-ovate, entire, somethat three-nerved at base;
cauline leaves linear, attenuated below;
external leaves of the involucrum longer than the inner, spreading, lanceolate, densely tomentose;
rays numerous, (twenty to twenty-four.)
Hab.
In the Rocky Mountains, by Flat-Head River, towards the sources of the Oregon.
Flower large and showy,
about three to four inches in diameter,
while the scapoid stem is not more than a span high.
Stigmas very hirsute, filiform.
Rays feminine, with infertile filaments.
- Elliott, Stephen, 1821-1824.
• Golden Checklist Flora:
Brickellia;
• Publication Details:
Elliott, 1821-1824, publication details;
Brickellia Elliot
The name is conserved against Brickellia Raf., Med. Repos. Ser. 2, 5:353 (1808), nom. Illeg. Nom rej., which turns out to be a synonym of Gilia Ruiz & Pav.
BRICKELLIA. E.
Involucrum polyphyllum, inbricatum.
Semina sub glabra, 10 striata.
Pappus pilosus sive scaber.
Receptaculum nudum, punctatum.
Involucrum many leaved, imbricate.
Seed nearly glabrous, 10 streaked.
Pappus hairy or scabrous.
Receptacle naked, dotted.
1. Cordifolia. E. …
…
This plant which in its artificial characters is closely allied to the Eupatorium,
differing principally in size and number, in its general aspect,
bear more resemblance to the Vernonia.
I have named it in commemoration of Dr. John Brickell, of Savannah,
who at one period of his life paid much attention to the botany of this country,
and made known to Dr. Muhlenberg, Fraser and others, many of its undescribed plants.
…
- Schilling, Edward, Randall W. Scott, and Jose L. Panero, 2015.
Key to the Sections of Brickellia
1 Cypselae with 4–6 ribs, sometimes obcompressed; plants annual or
perennial ...... 2
- Cypselae with 8–10 ribs, prismatic; plants
perennial ...... 3
2 Leaves setose, with prickles at tips of lobes; corollas
tubular .......................................................................... 1. B. sect. Barroetea
- Leaves not setose, lacking prickles at tips of lobes; corollas funnelform …… 8. B. sect. Phanerostylis
3 Inflorescence axis with few to numerous small, undeveloped heads (the crowded short phyllaries giving the appearance of gemmae)
interspersed with fully developed flowering heads …… 4. B. sect. Gemmipedium
- Inflorescence with all heads developing fully …… 4
4 Pappus bristles usually plumose; plants herbaceous perennials,
with usually alternate leaves and few-flowered (8–10) heads ……. 5. B. sect. Kuhnia
- Pappus not plumose, usually barbellate; plants herbaceous or shrubby, leaves alternate or opposite,
heads with few to many flowers …… 5
5 Larger leaves with naked basal veins,
the lateral veins marginal at the point of divergence from the midrib (Fig. 1) …… 2. B. sect. Brickellia
- Larger leaves with the first pair of lateral veins diverging from the midrib above the leaf margin …… 6
6 Plants annuals or weak perennials; heads 11–14 flowered; if perennial, leaves hastately toothed or lobed,
petioles 2–3.5 cm long …… 6. B. sect. Leptanthodium
- Plants perennials, herbaceous or shrubby; heads (3–)11–100-flowered;
if leaves hastately toothed or lobed, heads usually more
than 15-flowered or petioles less than 2 cm long …… 7
7 Leaves sessile or short-petiolate; plants usually shrubs; heads erect; corollas funnelform …… 7. B. sect. Microphyllae
- Leaves usually petiolate; plants shrubs or perennial herbs; heads erect or nodding;
corollas funnelform or tubular …… 8
8 Leaves usually cuneate at base; heads usually with 13–25 flowers;
involucral bracts usually weakly imbricate; pappus length
usually 6 mm or less; pappus bristles usually fewer than 40 …… 8. B. sect. Xerobrickellia
- Leaves usually truncate or cordate at base;
heads with (15)25–100 flowers; involucral bracts strongly imbricate; pappus length
usually 6 mm or more; pappus bristles usually more than 40 …… 3. B. sect. Coleosanthus
2. Brickellia sect. Brickellia.
- Schilling, Edward E., Jose L. Panero, Bonnie S. Crozier, Randall W. Scott, and Patricia Davila, 2015.
- Nesom, Guy L., 2022.
Key to USA species of Brickellia section Kuhnia After Neson (2022)
1. Florets 6–15 per head.
2. Outer and middle phyllaries often nearly equalling the inner in length, with apices long-attenuate
into falcate or twisted, filiform tips …… Brickellia ozarkana
2. Outer and middle phyllaries graduate in length, with apices apices appressed, acute to short-
acuminate, similar to the inner.
3. Leaves (all of the stem) linear, 6–18(–25) mm long, 1–2(–3) mm wide ……. Brickellia mosieri
3. Leaves (below the inflorescence) lanceolate-ovate to lanceolate or narrowly lanceolate, mostly
25–100 mm long, 5–40 mm wide, rarely linear as populational variants, if linear (as in var.
gracilis-Fig. 0 and Anderson 20487-Fig. 0) then 15–30 mm long, 2–3 mm wide
…… Brickellia eupatorioides sensu stricto
1. Florets (12–)14–35 per head.
4. Leaves linear; florets 15–35 per head …… Brickellia leptophylla
4. Leaves broader; florets (12–)14–24 (–29) per head
5. Outer and middle phyllaries often nearly equalling the inner in length, with apices long-
attenuate into loose and spreading, falcate or twisted, filiform tips …… Brickellia macranthra
5. Outer and middle phyllaries graduate in length, with apices appressed, acute to short-
acuminate, caveat …… Brickellia suaveolens
• Golden Checklist Flora:
Brickellia californica;
Brickellia californica (Torrey & A. Gray) A. Gray. “California Brickelbush”
- DeCandolle, Augustus Pyramus, 1836.
- Gray, Asa, 1849.
- Kunth, Karl Sigismund, 1833-1850.
• Publication Details:
Gray, 1849, publication details;
Original Text
Comments
308. B. (Bulbostylis) Californica, Torr. & Gray, Fl. 2. p. 79 :
var. foliis plerisque subcordatis.
—
Rocky hill-side on the Mora River, and eight miles eastward, in bottom land;
Aug. (346.)
—
The cauline leaves are mostly cordate,
and with rather longer petioles than in the Californian specimens collected by Douglas ;
but the plant of Hartweg's recent collection is wholly intermediate.
Mr. Bentham (in Bot. Voy. Sulph.) has very properly carried out the intimation given in the
Flora of North America, and annexed Bulbostylis to Brickellia.
The pappus is so strongly barbellate-denticulate in some species
(as in the original B. cordifolia and especially in B. cylindracea, Gray & Engelm.,
from Texas, and an undescribed Mexican species found by Dr. Wislizenus *)
that Clavigera is separated by a merely arbitary character.†
I almost fell off my chair to see that Torrey & Gray applied Bulbostylis DC. to our Brickellia.
DeCandolle (1836) did in fact propose it for a group in the Eupatorieae.
That name is now considered illegitimate
and the name in Cyperaceae, which would appear to be Bulbostylem Kunth, is a conserved name (Kunth, 1833-1850, v. 2, pg. 205).
- Schilling, Edward, Randall W. Scott, and Jose L. Panero, 2015.
• Golden Checklist Flora:
Brickellia eupatorioides;
Brickellia eupatorioides (L.) Shinners. “False Boneset”
- Linne, Carl von, and Lars Salvius, 1763.
Original Text
Eupatorioides.
289. KUHNIA. (Pentandra, Monogyna.)
Eupatoria conyzoides odorata, folio crenato molli fubincano. Pluk. alm. 140. t. 87. f. 2?
Habitat in Penfylvania , unde vivam attulit Adam Kuhn. ♃
Caules fesquioedales, erecti, laeves, rigiduli.
Folia alterna, petiolata, lato-lanceolata, nuda, fubrugefa, fubtus venofa, fusdentata ferraturis mediis majoribus.
Rami alterni, e medio caule.
Corymbi parvi, terminales & etiam fape ramorum lateralium.
Corollae albe.
Stamina albo-flavefcentia.
Planta refert Eupatorium, fed Piftilla clavata, &
Antherae diftinctae, cylindricaw, apice labio dehifcentes, absque exemplo in alio Compofits flore.
• Golden Checklist Flora:
Brickellia grandiflora;
Brickellia grandiflora (Hook.) Nutt. “Tasselflower Brickellbush”
- Hooker, Sir William Jackson, 1829-1840.
Original Text
COMPOSITAE Omissae
Post Eupatorium occidentale, v. 1. p. 305, adde
5.* Eupatorium? grandiflorum;
glabrum,
foliis alternis petiolatis cordato-triangularibus acuminatis grosse serratis venosis,
paniculae floribus glomeratis,
incolucri foliolis multiserialibus lineari-oblongis acutis striatis exterioribus e lata basi subulatis subsquarrosis,
acheniis cylindraceis striatis scabridis.
Hab.
On the low hills betweeb the north and south branches of Lewis and Clarke's River, in stony places.
Douglas.
—
Three to four feet high, herbaceous.
Stem rounded.
Flowers large, white, clustered.
The leaves bear a considerable resemblance to those of E. occidentale,
but the flowers are totally different, larger than is usual in this Genus,
each of many florets, and with the stigmas less exserted than usual.
I possess only a solitary specimen.
- Nuttall, Thomas, 1840-1841.
Original Text
Brickellia grandiflora,
leaves alternate, deltoid-cordate, acuminate, incisely dentate towards the base,
entire at the point, smooth on both surfaces, and covered beneath with resinous atoms;
flowers in fastigiate clusters, the upper part of the stem branching;
inner scales of the involucrum linear-lanceolate, acute;
pappus white, achenia smooth.
—
Eupatorium? grandiflorum.
Hook. Flor. Am., Vol. II., p. 26.
Hab.
In the Rocky Mountain range, by streams, in gravelly places, and west,
to the lower falls of the Columbia.
—
Perennial.
Stems many from the same root,
about twelve to fifteen inches high.
The whole plant almost perfectly glabrous.
Leaves alternate, sometimes almost opposite, approximate,
on longish petioles, deltoid-cordate, acuminate, coarsely and deeply toothed towards the base,
smooth and green, but shhining, aith a coating of yellow resinous atoms having a heavy aromatic scent;
stem branching above;
branches terminating in corymbulose clusters of subsessile flowers,
about five capituli in each.
Florets straw-yellow, inclining to white, cylindric and smooth,
the border connivent.
Stigmas exserted, smooth, thicker toward the extremity.
Achenium cylindric, ten-striate.
Pappus of a single series of twenty to twenty-four scabrous hairs.
Receptacle naked, flat.
Centaurea L “Knapweed”
- Hilpold, Andreas, et al., 2014.
The virtual absence of congruence between morphology and
molecular data suggests that the morphological characters that
have been used taxonomically are not reliable with respect to
the true relationships.
• Golden Checklist Flora:
Centaurea diffusa;
Centaurea diffusa Lam. “White Knapweed”
- Lamarck, Jean-Baptiste, 1783.
Original Text
Translation and Comments
70. Centauree diffufe ,
Centaurea diffufa.
Centaurea calycibus coliato-fpinulofis exiguis,
foliis radicalibus lyratis,
caulinis anguftis fubintefris ;
caule ramofiffimo paniculato diffufo.
N.
Carduus orientalis calcitrapae folio, flore monimo.
Tourn. Cor. 31.
70. Centauree diffuse
Carex diffused.
Carex ealycibus coliato-spinulosis small;
Radical leaves lyratis,
caulinis tightened subintefris;
Stem ramosissimo paniculato abroad.
X.
Eastern calcitrapae thistle leaf, flower Monimum.
Tournefort Cor. 31.
Cette plante reffemble beaucoup ŕ la Centaurée
panicuîee no. 36 , & pariot n'en différer efientîellement
que par les pointes de fes écailles calîcinales ,
qui font trčs-aiguës , roides & un peu
épineufes. La tige eft menue , anguleufe , légčrement
cotonneule , haute de prčs d'un pied, extręmeraent rameufe ,
panîculée , & diffure, Ses
feuilles inférieures font en lyre; toutes les autres
font petites , étroites , la plupart fimples , &
linéaires. Les fleurs font trčs-petites , extremement
nombreufes , & ŕ écailles calicinales-lancéolces ,
ciliées , & terminées par une pointe un peu épijieufe.
Cette plante croît dans le Levant, (v. f.)
This plant looks a lot like Knapweed
panicuîee no. 36, & pariot actually differ
than by the points of the calicinal scales,
which are very sharp, stiff & a little
thorns. The stem is slender, angular, slightly
cotton, nearly a foot high, extremely branched,
panicle, & diffure, Its
lower leaves are lyre-shaped; all the others
are small, narrow, mostly fimple, &
linear. The flowers are very small, extremely
numerous, with calicinal-lanceolate scales,
ciliate, & terminated by a somewhat epijieuve point.
This plant grows in the Levant.
(Seen in the dried state.)
• Golden Checklist Flora:
Centaurea stoebe;
Centaurea stoebe L. “Spotted Knapweed”
- Linne´, Carl von, 1753.
Original Text
Stśbe.
23 CENTAUREA calycibus ciliatis oblongis,
foliis pinnatifidis linearibus integerrimis.
Roy. lugdb. 140.
- Lamarck, Jean-Baptiste, 1783.
Original Text
Translation and Comments
35. Centauree mouchetee ,
Centaurea maculofa.
Centaurea calycibus ciliatis ovato-fubrotundis pulchre maculofis ,
foliis tenuibus bipinnatidifis , caule fubpaniculato.
N. Centaurea , no. 79 & 80. Gmel. Sib. 2. p. 99. Tab. 44. f. 1. 2.
Cette Centauree fe diftingue aifement de la
fuivante par fes feuilles decoupees tres-menu ,
& par fes fleurs , qui font au moins une fois plus
groffes , & ont leur calice agreablement mouchete.
Sa tige eft haute de huit a dix pouces ,
ftriee , blanchatre, & un peu paniculee vers fon
fommet. Ses feuilles font blanchatres . a decoupures
menues & pointues ; les inferieures font
oblongues & bipinnatifides ; les fuperieures font
plus petites , fimplement pinnees , & a decoupures
lineaires. Les fleurs font purpurines & remarquables
par leur calice gros, court, mouchete par
les taches bbrunes qui font au fomment de fes ecailles.
Nous avons obferve cette plante en Auvergne ,
aux environs de Clermont , fur le puits de Crouel.
( v.v. )
We have obtained this plant in Auvergne,
near Clermont, on Puy de Crouel (a small local hill).
Auvergne is a historical region in central France,
and is now part of Auvergne-Rhône-Alpes.
( I have seen it live. )
- Weber, William A., and Ronald C. Wittmann, 2012.
- Ackerfield, Jennifer, 2015.
I'd like to give credit and copyright information.
However, any attempt to find out more about CABI.org fails with a 504 Gateway Time-out.
Also it would be really helpful to find a copy of the Ochsmann (2001) paper
because many sources of information refer to it.
Original Text
Most of the North American plants called Centaurea maculosa are perennial, polycarpic and tetraploid (2n = 36).
However, there have been multiple introductions, and some infestations might be diploid.
Diploids can be identified by genetic analysis and growth habit,
because they flower once and die after 2 years (Smith, 2001).
Ochsmann (2001b) cites evidence of hybridization between spotted knapweed
and diffuse knapweed (Centaurea diffusa) in at least seven US states.
The hybrid is named Centaurea ×psammogena.
- Flora of North America Editorial Committee, eds., 1993+.
Original Text
Recent studies have shown that the American plants are identical with plants introduced
to the whole of Europe (J. Ochsmann 2001).
Subsp. micranthos, a tetraploid perennial, is clearly distinct from the diploid,
biennial plants native to central Europe known as C. stoebe Linnaeus subsp. stoebe,
C. rhenana Boreau, or C. maculosa Lamarck.
In most American literature the name Centaurea maculosa Lamarck has been misapplied to
C. stoebe subsp. micranthos.
W. A. Weber (1987, 1990) treated this taxon as Acosta maculosa (Lamarck) Holub.
The treatment of about 100 species of Centaurea sect. Acrolophus Cassini
as the genus Acosta by J. Holub (1972) and others is supported
by neither morphologic nor molecular characters and is not widely accepted in Europe.
- POWO, 2021 - 2026.
- Ackerfield, Jennifer, 2015.
Cirsium Mill. Thistle.
• Golden Checklist Flora:
Cirsium ochrocentrum;
Cirsium ochrocentrum A. Gray “Yellowspine Thistle”
- Flora of North America Editorial Committee, eds., 1993+.
• Golden Checklist Flora:
Cirsium undulatum;
Cirsium undulatum (Nutt.) Spreng. “Wavy Leaved Thistle”
Cirsium undulatum is widely distributed in the wstern half of North America
from the dry plains and plateaus of the Pacific Northwest eastward across the Great Plains
to Manitoba and the Dakotas and south to Texas, New Mexico, and northwestern Mexico.
It occurs in scattered localities in the Rocky Mountains and northeastern Great Basin region.
At least some of the few widely scattered records from the eastern United States are probably introductions.
Cirsium undulatum is both widespread and variable.
Plants of the Great Plains region tend to be low-growing with a few large heads and elongate corollas.
Plants of the Pacific Northwest are usually taller and produce smaller, more numerous heads with shorter corollas.
A detailed study of this species might reveal races worthy of recognition as infraspecific taxa.
— David J. Keil in Flora of North America (vol. 19, 20 and 21)

Distribution of collections of Cirsium undulatum
- Miller, Philip, 1754.
- Nuttall, Thomas, 1818.
• Glossary:
ventricose;
• Publication Details:
Nuttall, 1818, publication details;
538. CARDUUS. L. (Thistle.)
Calix ventricose, imbricate, scales spiny.
Receptacle villous.
Pappus pilose or plumose, deciduous.
§ II. CNICUS. Pappus plimose
11. * undulatus.
Stem low and few-flowered;
leaves amplexicaule, pinnatifidly sinuate, and plicately undulated, on both sides tomentose, but beneath white, lobes bifid and spiny;
calix subglobose, scales lanceolate, eredt and mucronate.
HAB.
On the calcareous islands of lake Huron, and on the plains of Upper Louisiana.
—
Leaves almost like some species of Cynara, but not remarkably large, moreslenderly tomentose on the upper side;
stem often 1,2, or few-flowered, and 1 to 2 feet high.
Flowers large, reddish purple.
- Sprengel, Curt Polycarp Joachim, 1826.
Original Text
Translation
undulatum
*
45. C. caule humili paucifloro,
foliis amplexicaulibus finuato-pinnatifidis undulato-plicatus utrinque tomentofis,
laciniis 2fidis fpinofis,
fquamis anthodii lanceolatis erectis mucronatis.
In infulis lacus Huronum et Louifiana fuper.
(Cnicus Nutt.)
undulatum
*
Cirsium [with] short few-flowered stems,
leaves amplexicaul curved-pinnatifid undulate-plicate both sides hairy,
lobes divided spiny scales anthodii laneeolatis putting pointed.
The islands of Lake Huron and upper Louisiana.
- Frankton, C., and R. J. Moore, 1961.
Abstract.
The morphology and specific differences of Cirsium undulatum (Nutt.) Spreng. And of C. flodmanii (Rydb.) Arthur
are described and their Canadian distributions are reported in detail.
The chromosome numbers are C. undulatum f. undulatum and f. album Farwell, 2n = 26;
C. flodmanii f. flodmanii and f. albiflorum D. Löve, 2n = 22.
The origin of four North American species of Cirsium that do not follow the world-wide base number 17 is discussed;
it is postulated that reduction in number has occurred by translocations.
The chromosomes of species with reduced numbers are larger than those of the unreduced species
but the total length of the chromosomes of both groups is approximately the same.
- Häffner, Eva, and Frank H. Hellwig, 1999.
- Garcia-Jacas, Nuria, Teresa Garnatje, Alfoonso Susanna, and Rosier Vilatersana, 2002.
- Kelch, Dean G., and Bruce G. Baldwin, 2003.
- Bodo Slotto, Tracey A., David P. Horvath, and Michael E. Foley, 2012.
Abstract. Weedy invasive Cirsium spp. are widespread in temperate regions of North America and some of their biological control agents have attacked native Cirsium spp. A phylogenetic tree was developed from DNA sequences for the internal transcribed spacer and external transcribed spacer regions from native and non-native Great Plains Cirsium spp. And other thistles to determine if host specificity follows phylogeny. The monophyly of Cirsium spp. And Carduus within the tribe Cardinae was confirmed with native North American and European lineages of the Cirsium spp. Examined. We did not detect interspecific hybridization between the introduced invasive and the native North American Cirsium spp. Selected host-biological control agent interactions were mapped onto the phylogenic tree derived by maximum likelihood analysis to examine the co-occurrence of known hosts with biological control agents. Within Cirsium-Cardueae, the insect biological control agents do not associate with host phylogenetic lines. Thus, more comprehensive testing of species in host-specificity trials, rather than relying on a single representative of a given clade may be necessary; because the assumption that host-specificity follows phylogeny does not necessarily hold. Since the assumption does not always hold, it will also be important to evaluate ecological factors to provide better cues for host specificity.
• Golden Checklist Flora:
Cirsium vulgare;
Cirsium vulgare (Savi) Ten. “Bull Thistle”
• Golden Checklist Flora:
Conyza canadensis;
Conyza canadensis L.
• Golden Checklist Flora:
Coreopsis tinctoria;
Coreopsis tinctoria Nutt. “Golden Tickseed”
- Nuttall, Thomas, 1821.
• Publication Details:
Nuttall, 1821, publication details;
Original Text
1. COREOPSIS * tinctoria,
foliis radicalibus pseudobipinnatis,
foliolis subovalibus integris glabris,
superioribus pseudopinnatis laciniis linearibus;
floribus binatis ternatisve;
calcibus exterioribus brevissimis;
radiis bicoloribus;
seminibus nudis immarginatis.
Habitat.
Throughout the Arkansa territory to the banks of Red River,
chiefly in the prairies which are subject to temporary inundation.
—
Flowering, from June to October.
Description.
Annual and biennial, stem erect, smooth, and much branched,
extremely variable in magnitude,
being from one to five feet high.
The leaves, in common with the genus,
are somewhat think and succulent,
the primary ones simple,
radical pseudobipinnate, the segments also occasionally pinnate,
oblong-oval, commonly smooth, and entire,
the ultimate divisions largest.
Flowers often terminating the branchlets by pairs,
with the peduncles unusually short.
Exterior calix, minute, much shorter than the interior,
and in common with it.
And the number of rays mostly eight-leaved.
Rays three-lobed at the extremity,
of a bright orpiment yellow and brown towards the base;
disk brown, and rather small.
Receptable paleaceous, the leaflets deciduous.
Seed small, blackish, immarginate,
curved and naked at the summit.
Economical Use.
The flowers of this species afford a yellow dye,
in common with those of the C. senifolia.
As an ornamental plant, of easy culture and uncommon brilliance,
it promises to become the favourite
of every garden where it is introduced.
• Golden Checklist Flora:
Crepis occidentalis;
• Publication Details:
Nuttall, 1834a, publication details;
Crepis occidentalis Nutt. “Largeflower Hawksbeard”
- Nuttall, Thomas, 1834a.
COMPOSITAE
50. Crepis *Occidentalis.
Canescente-pubescens, pumila, foliis sessilibus,
runcinato-pinnatisentis, laciniis linearibus,
acutis subdenticulatis, floribus paucis fastigiatis.
♃
About a span high, covered with a close very short whitish pubescence,
The leaves runcinate and acute,
about two on the spem,
greatly resembling those of the common Shepard's purse ;
above, beneath the ultimate flowers,
diminishing into simple undivided bracts.
Flowers (in the only specimen before me) three,
axillary and terminal,
all attaining nearly the same height on the stem.
Calyx slightly caliculate,
the larger leaves of it disposed in a single series,
the divisions linear and rather obtuse.
Flowers bright yellow, rather large,
about the size and appearance of those of Apargia autumnalis ;
liguli five-toothed ;
the anthers simple, the style bifid,
deeply and far exserted.
Pappus pilose,
somewhat scabrous through a lens,
the hairs more than twenty,
not dilated at base,
or in any way distinguishable from those of Hieracium,
nor are they all stipitate.
Seed smooth, brown.
Probably the Hymenonema laciniatum of Hooker,
in Flor. Boreal. Amer. L. c.
Hab.
Common on the borders and in the vicinity of the river Columbia.
- Nuttall, Thomas, 1840-1841.
• Glossary:
achenium;
testa;
• Publication Details:
Nuttall, 1841, publication details;
Original Text
* PSILOCHENIA
Crepis,
but with the achenium cylindric, curved, narrower above,
and without any visible striae, the testa indurated, and, when mature, black ;
an abortive outer series of florets, with the achenium empty.
Pappus copious, slenderly pilose, scabrous, and yellowish white,
about the length of the achenium.
Receptable naked, alveolate, the alveoles minutely fringed.
—
A low perennial herb; stem dichotomous and corymbose.
Leaves lanceolate, runcinately pinnatifid,
and, as well as the somewhat hirsute involucrum,
cinereously and closely lanuginous;
flowers yellow, rather large.
Psilochenia * occidentalis. Crepis occidentalis, Nutt.
In Journ. Acad. Nat. Sci. Philad., Vol. VII., p. 29.
Hab.
On the plains of the Platte, towards the Rocky Mountains.
The whole plant more or less canescently pubescent.
Stem about six or seven inches high, forked and corymbose at the summit.
Leaves about an inch wide, four or five inches long,
deeply and runcinately pinnatifid,
the segments linear-lanceolate and denticulate, uppermost leaves linear.
Involucrum campanulate;
sepals about twelve to fifteen in a single series, linear and somewhat acute;
involucel or bractes four or five, small and subulate:
there are blackish hairs mixed with the hoary pubescence of the sepals.
Florets about twelve, yellow, exserted.
- Babcock, E. B., and G. Ledyard Stebbins, 1938.
Original Text
TAXONOMIC HISTORY OF THE INDIGENOUS SPECIES
The first record of the occurrence of Crepis in North America
(outside of the arctic regions)
was made by Hooker in his “Flora Boreali-Americani” (1834, vol. 1, p. 297),
in which he identified as the European C. biennis James's Hieracium runcinatum.
In the same year Nuttall published his C. occidentalis,
and seven years later (1841),
in describing the plants collected on a tour to the Pacific
added another species, C. acuminata.
In this publication he created a new genus, Psilochenia,
for C. occidentalis,
and another, Crepidium,
for Hieracium runcinatum (Hooker's “C. biennis” of America),
and placed Crepis acuminata in a new sub-genus, Leptotheca.
Two new species, Crepidium glaucum and C. caulescens, were also described.
Two years later Torrey and Gray (1843) reduced Nuttall's two genera to Crepis,
and recognized, in addition to the arctic-alpine C. nana Richards.,
and C. elegans Hook., four species,
C. runcinata, C. glauca, C. occidentalis, and C. acuminata.
For the next fifty-three years the group received little attention,
although a few species and varieties were described by Gray and others.
In his “Syntopical Flora of North America” (1876) Gray recognized,
in addition to the six species included in his earlier work, two more,
C. Andersonii Gray and C. intermedia Gray,
and listed two varieties under the latter and three under C.occidentalis.
With the heading of the section including C. occidentalis, intermedia,
and acuminata he made the terse comment “species difficult,”
an opinion with which all later workers on the group,
including the present writers, are inclined to agree.
Coville (1896) was the first to make a monographic study of any of the American species.
Omitting the arctic-alpine species and the group of C. runcinata,
as well as C. acuminata and C. intermedia,
which latter he considered to be not well enough understood,
he included seven species, four of which,
C. monticola, C. scopulorum, C. rostrata, and C. barbigera,
were described as new.
His treatment, although well worked out according to the knowledge then available,
was based on a relatively meager series of specimens,
and included little information on the interrelationships of the species,
while their distribution could be stated only in relatively broad terms.
Since the work of Coville,
no particular attention has been paid to the genus in North America,
except for the description of numerous new “species,”
mostly of the group of C. runcinata,
by Greene, Rydberg, and others.
These were published without consideration of the group as a whole,
were based on fickle characters such as leaf shape and pubescence,
and none of them are considered valid by the present writers.
Meanwhile in the affinity of C. occidentalis and
C. acuminata the species were variously interpreted by the writers
of the different floras, the keys to them varied considerably,
while the identification of the ever increasing number of collections
by various students of the western flora bore out Gray's opinion,
“species difficult.”
- Weber, William A., 1983.
Weber then goes to to propose 23 new combinations in Psilochenia.
Original Text
The American species of Crepis were treated in a now classic
monograph by Babcock & Stebbins (Carnegie Inst. Wash. Publ. 504. 1938).
The authors seem to have been preoccupied with the species alone,
and unfortunately they did not discuss the significance of
their cytological findings as having a bearing on the generic level,
even though Nuttall (1841) had proposed the name
Psilochenia for the American species.
All of the native American species of Crepis,
with the exception of two Old World species (C. elegans and C. nana),
representing an ancient Tertiary extension of the genus onto western North America,
have the chromosome base number x=11.
“This is in striking contrast to the Old World species of Crepis,
whose basic haploid numbers range from x=3 to x=7,
4 and 5 being much the most common" (Babcock & Stebbins, op. cit.).
The authors went on to postulate that the American species
may have arisen by amphidiploidy from a cross involving
Crepis species with x=4 and x=7.
Whether or not this can ever be substantiated,
the fact remains that the American species of Crepis
form an indisputably monophyletic line,
spatially and genetically isolated from the Old World species.
Recently Love (1982, p. 360) transferred Crepis runcinata
to Nuttall's genus Psilochenia because of this evidence.
but among the rest of the species, only the type,
Psilochenia occidentalis Nuttall, has a name in that genus.
The following combinations are needed.
- Enke, Neela, 2009.
• Taxa Notes:
Crepis occidentalis, Weber & Wittman, 2012;
Original Text
Crepis acuminata differs from all other species of clade V in morphology, karyology and geographic distribution.
As all members of sect. Psilochaenia it occurs exclusively in North America,
is polyploid and has a basic chromosome number of x=11.
The singularity of these features within Crepis s.str. Support the maintenance of sect. Psilochaenia.
Original Text
Crepis sect. Psilochenia (Nutt.) Babc. (1947) ≡ Psilochenia Nutt. (1841).
— Type: C. occidentalis Nutt.
?C. atribarba A. Heller
?C. bakeri Greene
?C. barbigera Coville
?C. intermedia A. Gray
?C. modocensis Greene
?C. monticola Coville
?C. occidentalis Nutt.
?C. pleurocarpa A. Gray
?C. runcinata (E. James) Torr. & A. Gray
- Babcock, E. B., and G. Ledyard Stebbins, 1938.
- Weber, William A., 1983.
- Weber, William A., and Ronald C. Wittmann, 2012.
• Taxa Notes:
Crepis occidentalis, Enke, 2009;
Solidago nana, Nuttall, 1841;
Original Text
The genus Psilochenia encompasses the indigenous North American species of Crepis
(with the exception of the Old World taxa C. elegans and C. nana,
which represent a Tertiary extension).
The chromosome base number x=11 is unknown in Eruasia.
See Babcock & Stebbins 1938, Weber 1983.
The FNA ignores Psilochenia altogether.
- Ackerfield, Jennifer, 2015.
• Golden Checklist Flora:
Cyclachaena xanthifolia;
Cyclachaena xanthifolia (Nutt.) Fresen. “Carelessweed”
- Nuttall, Thomas, 1818.
• Publication Details:
Nuttall, 1818, publication details;
602. IVA. L.
Calix about 5-leaved, or 5-parted.
Feminine florets of the ray 5, naked.
Receptacle setosely paleaceous.
Seed obovate, naked.
Herbaceous or shrubby;
leaves 3-nerved, mostly carneous, opposite and alternate;
flowers spiked or paniculated, axillar and terminal.
Species.
1. ciliata. ...
2. * xanthiifolia.
Annual;
leaves opposite, petiolate, cordate-ovate, acuminated, douply serrate,
softly villous, beneath canescent;
spikes paniculated, naked;
calix 5-cleft.
Hab.
In arid soils, near Fort Mandan, &c. on the banks of the Missouri.
Obs.
Plant very large, 5 or 6 feet high,
with leaves nearly of the size and form of Xanthium Strumarium,
but covered with a soft and almost velvet-like villus;
upper leaves ovate;
flowers extremely numerous, in a diffuse panicle.
Calix 5-cleft, divisions ovate-lanceolate, acuminate.
Style of the discal florets simple;
stigma subcapitate.
Fertile florets 5, naked.
Receptacle subsetaceous.
—
Flowering in August.
3. imbricata. ... 4. axillaris. ... &c.
A North American genus with the exception of I. annua
indigenous to the tropical regions of the same continent.
- Fresenius, Georg, 1836.
• Publication Details:
Fresenius, 1836, publication details;
41.4. If, for a name of a genus or taxon of lower rank published before
1 January 1953, no reference to a basionym is given but the conditions for
its valid publication as the name of a new taxon or replacement name are
fulfilled, that name is nevertheless treated as a new combination or name
at new rank when this was the author’s presumed intent and a potential
basionym (Art. 6.10) applying to the same taxon exists.

Footnotes from Index Seminum in which Cyclachaena is proposed
Page 1 from Index Seminum Frankfort/Am Main
- Miao, Bomao, Billie L. Turner, and Tom Mabry, 1995.
- Flann, C (ed), 2009+.
Dieteria Nutt.
- Nuttall, Thomas, 1840-1841.
Dieteria Nuttall
*DIETERIA.
Flower radiate, rays styliferous, fertile? liguli one or two series, broadish, those
of the disk hermaphrodite, fertile. Stigma filiform, hirsute and exserted.
Sepals of the involucrum, for the most part, closely imbricated in two to four
series, scariose and carinate, the tips usually reflected and herbaceous.
Receptacle flat or convex, alveolate, the alveolae deep, with toothed and
lacerated margins. Achenium obovate, subcylindric, ten to fifteen striate, pubescent.
Pappus of several series, scabrous and unequal, that of the ray shorter
and less copious.
—
Annual or biennial, (in one anomalous species perennial,)
divaricately branching herbs, more or less pubescent; leaves nearly entire,
incisely serrate or pinnatifid, the points often pungently mucronulate.
Flowers fastigiate. The disk yellow. Liguli red or purple.
—
Allied to Aster,
but with the involucrum regular; the achenia convex, distinctly striate
when ripe ; the receptacle deeply alveolate ; the pappus of the ray different
from that of the disk; the leaves incise or pinnatifid, and the duration only
to the first period of flowering. They are also allied to the first section of
Heterotheca by the deficient pappus of the ray, but that of the disk is simple,
and the rays are purple. The whole plant bitter to the taste.
—
(So called from their biennial duration.)
- Nuttall, Thomas, 1840-1841.
Original Text
* DIETERIA.
Flower radiate, rays styliferous, fertile ? liguli one or two series, boradish, those of the disk hermaphrodite, fertile.
Stigma filiform, hirsute and exserted.
Sepals of the involucrum, for the most part, closely imbricated in two to four series, scariose and carinate, the tips usually reflected and herbaceous.
Receptacle flat or convex, alveolate, the alveolć deep, with toothed and lacerated margins.
Achenium obovate, subcylindric, ten to fifteen striate, pubescent.
Pappus of several series, scabrous and unequal, that of the ray shorter and less copious.
—
Annual or biennial, (in one anomalous species perennial,)
divaricately branching herbs, more or less pubescent;
leaves nearly entire, incisely serrate or pinnatifid, the points often pungently mucronulate.
Flowers fastigiate.
The disk yellow.
Liguli red or purple.
—
Allied to Aster, but with the involucrum regular;
the achenia convex, distinctly striate when ripe;
the receptacle deeply alveolate;
the pappus of the ray different from that of the disk;
the leaves incise or pinnatifid, and the duration only to the first period of flowering.
They are also allied to te first section of Heterotheca
by the deficient pappus of the ray, but that of the disk is simple,
and the rays are purple.
The while olant bitter to the taste.
—
(So called from their biennial duration.)
• Golden Checklist Flora:
Dieteria bigelovii;
Dieteria bigelovii (A. Gray) D. R. Morgan & R. L. Hartman “Bigelow's Tansy Aster”
- Torrey, John, 1857.
Original Text
Aster Bigelovii (sp. nov,) :
ramis viscido-hirsutis ad apicem usque foiiosis; ramulis corymbosis monocephalis ;
foliis membranaceis oblongo-lanceolatis semiamplexicaulibus grosseserratis
fenuiter triplinerviis hirto-puberulis glabratis ; capitulis magnis globosis ;
involucri pluriserialis squamis attenuato-subulatis basi appressis superne longe caudato-appendiculatis
squarroso-recurvis glanduloso-viscidis ; acheniis glaberrimis.
Arroyos in the Sandia mountains ;
October.
A wholly new and most remarkable Aster, of the Grandiflori group ; but the apparently
showy heads larger than those of A. grandiflorus, being an inch in diameter, and the numerous
(blue and violet) rays an inch long. It is probably a tall plant ;
but the base of the stem was not collected.
Cauline leaves two or three inches long, coarsely dentate-serrate throughout ;
the uppermost, and those of the short branchlets, smaller and less toothed.
Scales of the imbricated involucre half an inch long when extended, very slender ;
the long and almost filiform appendicular portion recurved, spreading and very glandular.
Receptacle flat, alveolate ; the alveolae short and entire.
Achenia perfectly glabrous, linear, compressed, three lines long.
Pappus not abundant, nearly in a single series.
• Golden Checklist Flora:
Dieteria bigelovii bigelovii;
Dieteria bigelovii (A. Gray) D. R. Morgan & R. L. Hartman var. bigelovii “Bigelow's Tansy Aster”
• Golden Checklist Flora:
Dieteria canescens;
Dieteria canescens (Pursh) Nutt. “Hoary Tansyaster”
- Pursh, Frederick, 1814.
Original Text
636. ASTER. Gen. pl. 1291.
* Foliis integerrimis.>
…
17. A. cano-pubescens ;
foliis linearibus, panicula corymbosa ramisissima foliosa,
calycibus imbricatis acutissimus disco longioribus.
canescens.
On the banks of the Missouri.
♄.
Aug.-Nov.
v. s.
Flowers the size of a daisy, rays pale purple.
Seen by Pursh in the dried state, but neither the collector nor the herbarium is identified.
Probably was not Lewis & Clark, because their single specimen was collected on the Columbia River and first thought to be Aplopappus sp.
More likely a Nuttall collection, though Bradbury cannot be ruled out.
- Nuttall, Thomas, 1840-1841.
Original Text
† Involucrum subovate, of three or four series of scales.
Dieteria canescens;
leaves entire, linear, sessile, radical spathulate;
stem low and much branched, canescently villous, as well as the involucrum;
flowers fastigiate;
rays about eighteen to twenty; pappus very slender.
Hab.
On the denuded banks of the Missouri.
Aster canescens, Pursh, Bor. Am., Vol. II., p. 547.
Not in the least allied to Aster multiflorus.
A. biennis, Nutt. Gen. Am., Vol.II., p. 155.
I doubt if the leaves are always entire,
a fact so contrary to all the rest of the genus to which it is,
in all other respects, so intimately allied.
• Golden Checklist Flora:
Dyssodia papposa;
Dyssodia papposa (Vent.) Hitchc. “Fetid Marigold”
- Ventenat, Etienne P., Jacques M. Cels, and Henri J. Redoute, 1801.
- Hitchcock, A. S., 1891.
Original Text
• How did rubber rabbitbrush get that long scientific name?:
Introduction;
Ericameria nauseosa (Pall. ex Pursh) G.L.Nesom & G.I.Baird var. graveolens (Nutt.) Reveal & Schuyler
• Golden Checklist Flora:
Ericameria nauseosa nauseosa;
Ericameria nauseosa (Pall. ex Pursh) G. L. Nesom & G. I. Baird var. nauseosa. “Rubber Rabbitbush”

Habitat of Coll. No. 2440, Ericameria nauseosa var. nauseosa
Coll. No. 2440, Ericameria nauseosa var. nauseosa
- Hall, Harvey M., and Frederick E. Clements, 1923.

Hall & Clements (1923) Plate 34 of Ericameria nauseosa nauseosa.
• Notes on Colorado Flora:
Ericameria parryi;
Ericameria parryi (A. Gray) G. L. Nesom & G. I. Baird “Parry's Rabbitbrush”

Distribution of vouchers of Ericameria parryi in Colorado.
- Ackerfield, Jennifer, 2022.
- Urbatsch, Lowell E., et al., 2006.
- Weber, William A., and Ronald C. Wittmann, 2012.
Key to Varieties of Ericameria parryi
1
Distalmost leaves overtopping arrays, leaves sometimes gland-dotted.
2
1'
Distalmost leaves usually shorter than or equaling (seldom overtopping) arrays,
leaves not gland-dotted.
4
2
Leaves minutely stipitate-glandular.
Ericameria parryi var. parryi
2'
Leaf faces glabrous, tomentulose, tomentose, or moderately hairy, sometimes gland-dotted.
3
3
Leaves 30-80 mm. × 2-3 mm. wide;
leaf faces glabrous or puberulent, often minutely stipitate-glandular;
florets 8–20, corollas yellow.
Ericameria parryi var. parryi
3'
Leaves 20-40 mm. × 1 mm. wide;
leaf faces tomentose, eglandular;
florets 5–7(–8) corollas pale yellow;
n Arizona, Colorado, Nebraska, New Mexico, s Utah, Wyoming
Ericameria parryi var. howardii
4
Leaf faces tomentulose;
corolla pale yellow, tubes glabrous or puberulent, throats abruptly dilated, lobes 0.7–1 mm,;
Ariz., [Colo,] N.Mex., Utah.
Ericameria parryi var. affinis
4'
Leaves glabrous or sparsely hairy, sometimes ± viscid;
Corolla clear yellow, tubes glabrous, lobes 1.5–2 mm., throats gradually dilated, lobes 1.5–2 mm;
Colorado, New Mexico, Utah
Ericameria parryi var. attenuata
- Gray, Asa, 1863.
• Taxa Notes:
Notes on Ericameria parryi parryi;
...
*
293 (and 413 of Parry,) L. (Chrysothamnus) Parryi, n. sp.
†
A very distinct
species, which is said to abound in the Middle Park, South Park, and all
that district; the wonder is that it has not been detected before.
The spiciform or racemose and leafy inflorescence,
and the large heads with lax and taper-pointed scales, are characteristic.
...
†
Linosyris (Chrysothamnus) Parryi (sp. nov.):
fruticosa; ramis virgatis lanoso-dealbatis ;
foliis linearibus fere glabris subviscosis, floralibus conformibus capitula in thyrsum angustum
congesta longe superantibus ; involucro 10-15-floro cylindraceo pauciseriali, squamis sublaxe
imbricatis albidis lanceolatis, omnibus (exterioribus saepius folioso-interioribus scarioso-) attenuato-
acuminatis ; corollae tubo hirsutulo; acheniis linearibus cano-pubescentibus. Leaves 2 to 3
inches long, 3-nerved, acute, plane, the larger ones 2 lines wide and tapering to the base. Thyrsus
narrow, often almost simply racemose or spiciform, sometimes more compound and branchy. Heads
about two-thirds of an inch long, foliose-bracteate; the bracts passing into the exterior and leafy-
tipped scales of the involucre. Receptacle, styles, viscidity, aroma, &c., as in Chrysothamnus
generally.
- Greene, Edward Lee, 1895.
20. C. Parryi.
Linosyris Parryi, A. Gray, Proc. Philad. Acad. 66 (1863).
Bigelovia Parryi, A. Gray, Proc. Am. Acad. l. c.
Aster Harbourii, O. Ktze, 1. c. 316.
Stout, a foot high or more, quite leafy, and with ascending branches:
leaves linear, narrowed at base, 2 inches long or more,
somewhat tomentose like the branches, or glabrate: heads quite
thyrsoidly disposed up and down the branches, large, 10 to
15-flowered; involucral bracts lanceolate, attenuate-acuminate:
corolla-tube hirsutulous.
Parks of the Colorado Rocky Mountains.
- Nelson, Aven, 1899.
Chrysothamnus Parryi (Gray) Greene, l. c.
—
The habitat and range of this well-known species is always given as “parks of the
Rocky mountains in Colorado,” but certainly similar parks in
southern Wyoming, at least, must be added. Typical specimens
are, J. H. Cowen, Breckinridge, Colo., August 1896; the writer's
Nos. 2617 and 3495, Lincoln gulch, August 1896 and Big creek,
August 1897.
- Anderson, Loran C., 1984.
- Anderson, Loran C., 1995.
- Urbatsch, Lowell E., et al., 2006.
• Notes on Colorado Flora:
Ericameria parryi affinis;
Ericameria parryi var. affinis “Parry's Rabbitbrush”
Chrysothamnus affinis, n. sp.
—
Scarcely shrubby, the persistent
base hardly more than a much-branched woody caudex: the season’s
stems very numerous, simple, 1-2" high, yellowish, glabrate:
leaves crowded, narrowly linear, acute, erect or spreading, dark
green, nearly glabrous, 3-4cm long: inflorescence a crowded
spicate thyrsus which at maturity distinctly surpasses the leaves :
bracts glabrate, arachnoid-ciliate on the margins, somewhat
thickened-coriaceous, about three in each row; the outer with an
ovate base, contracted in a usually spreading acumination ; the
inner linear-oblong, abruptly acuminate, shorter than the pappus:
corolla tube slender, bearing only a few, minute, scattering clavellate
hairs, shorter than the expanded, tubular throat which 1s
cleft about one fourth its length: style appendages tardily but
at length wholly exserted: akene linear-cylindric, about 6mm long.
Allied to C. Howardi (Parry) Greene, but clearly distinct by its seat
shrubby habit, its greenish glabrate aspect, and its crowded yellow inflorescence
which surpasses the leaves. C. Howardi has cinerous leaves and
stems, a dirty whitish-yellow inflorescence, and the leaves overtop the
comparatively few and large heads whose bracts are distinctly arachnoid.
Excellent specimens were collected by J. H. Cowen, Jefferson, Colo.,
August 1896, and distributed as Bigelovia Howardi Gray. Type in Herb.
University of Wyoming.
Ericameria parryi var. attenuata “Parry's Rabbitbrush”
- Nelson, Aven, 1899.
Chrysothamnus affinis attenuatus, n. comb.
(Bigelovia Howardi attenuata Jones, Proc. Cal. Acad. Sci. II. 5:691: 1895.
—
The specimen of Mr. Jones’ type number, 5912, seems to
approach C. affinis much more closely than it does C. Howardi. Its
more persistent stems and longer branchlets, its very 1ong acuminate
bracts, and long exserted styles will readily distinguish it.
Ericameria parryi var. howardii “Parry's Rabbitbrush”
- Gray, Asa, 1865.
Linosyris (Chrysothamnus) Howardii, Parry, in litt. :
fruticosa,
semi-sesquipedalis ; ramis junioribus lanoso-dealbatis ; foliis linearibus
acutatis uninerviis glabratis (primum tenuiter arachnoideis), floralibus
capitula corymboso-congesta superantibus vel subeequantibus ; involucro
5-floro cylindraceo, squamis laxiusculis lanceolatis omnibus modo
L. Parryi attenuato-acuminatis ; corollas pallide flavoe tubo parce villosulo ;
acheniis linearibus pubescentibus. Var. α. foliis angustissime
linearibus elongatis ; involucro tenuissime arachnoideo glabrato.
—
Colorado Territory, on gravelly hills near “Hot Springs” of Middle
Park, where it is often associated with L. Parryi, Gray, coll. C. C.
Parry, 1863. It is more shrubby than that species, its smaller and
fewer-flowered heads in terminal, fastigiate, almost capitate clusters,
&c. A little of it was included under Hall and Harbour's No. 293.
Var. β. Nevadensis : foliis pi. m. latioribus brevioribus
(inferioribus subspathulato-linearibus), apice cuspidate saspius incurvo ;
involucri squamis prsesertim ad margines magis arachnoideis subviscosis.
—
Mount Davidson, Nevada, above Virginia City, H. G. Bloomer.
Near Carson City, Dr. C. L. Anderson.
Ebbett's Pass, within the borders of California, alt. 9,000 feet.
Prof. Brewer ; a dwarf form, only 5 or 6 inches high.
- Greene, Edward Lee, 1895.
21. C. Howardii.
Linosyris Howardii, Parry in Gray, Proc. Am. Acad. vi. 541 (1865).
Bigelovia Howardii, A. Gray, l. c. viii. 641 (1873).
Aster Howardii, O. Ktze, l. c. 318 (1891).
Chondrophora Howardii, Britton, Mem. Torr. Club, v. 317 (1894).
Similar to the last [C. parryi] in habit, rather
smaller: leaves linear, firm: heads 5-flowered, the 12 to 15
bracts of the involucre oblong-lanceolate, somewhat arachnoid,
the outer gradually, the inner abruptly cuspidate-acuminate:
corolla-tube sparingly villous.
Parks of the Colorado Rocky Mountains; but included by
Dr. Britton in the list of plants of the Gray’s Manual region,
I know not upon what grounds.
- Nelson, Aven, 1899.
Chrysothamnus Howardi (Parry) Greene,3 l.c.
—
The habitat of this, like the preceding, is given as “parks of the Rocky mountains,”
but it should be stated that the parks are very different
in their character, C. Parryi inhabits moist open ground, known
as parks, occurring at intervals in the timbered mountain ranges.
C. Howardi inhabits that other class of parks, viz., extensive,
high, dry table-lands like North Park, Colo, and the Laramie
Plains. It occupies the dry foothills and ridges and I doubt not
may occur on the eastern base of the Rockies, possibiy extending
into eastern Nebraska, as given in Britton & Brown’s Flora,
a statement questioned by Dr. Greene. It seems probable that
some confusion on this point has arisen by the distribution of a
somewhat similar plant occurring in situations much like those
in which C. Parryi occurs. This plant I think should stand as a
species, and may be named as below.
3Dr. Greene’s papers cite the literature of all the well-known species so fully
it seems necessary only to call attention to this fact.
Ericameria parryi var. nevadensis “Parry's Rabbitbrush”
22. C. Nevadensis.
Linosyris Howardii, var. Nevadensis, A. Gray, Proc. Am. Acad, vi. 541 (1865).
Bigelovia Nevadensis, A. Gray, Syn. Fl. 136 (1884).
Taller than the last, the head more elongated, and bracts more numerous,
hirsute-ciliate, all with long slender firm but recurved tips.
Eastern slope of the Sierra Nevada in California, and
adjacent Nevada.
- Gray, Asa, 1863.
• Notes on Colorado Flora:
Ericameria parryi parryi;
• Taxa Notes:
Notes on Ericameria parryi, Gray, 1863;
Ericameria parryi (A. Gray) G. L. Nesom & G. I. Baird var. parryi “Parry's Rabbitbrush”
• Golden Checklist Flora:
Erigeron canadensis;
Erigeron canadensis (L.) Cronquist “Canadian Horseweed”
- Linne´, Carl von, 1753.
3. ERIGERON caule floribusque paniculatis.
Hort. cliff. 407.
Hort. upf. 258
Gron. virg 96.
Gort. gelr 475
canadenfe.
Conyza annua acris alba elatior, linariae foliis.
Morif. hift. 3. p. 115. f. 11. t. 20. f. 29.
Bocc. fic. 85. t. 46.
Habitat in Canada, Virginia, nunc in Europa auftrali.
☉
- Nesom, Guy L., 2008.
Erigeron L. “Fleabane; Daisy”
• Golden Checklist Flora:
Erigeron compositus;
Erigeron compositus Pursh “Cutleaf Daisy”
- Moulton, Gary E., 1999.
- Pursh, Frederick, 1814.
Syngenesia Superflua. Erigeron
535
14.
E. pilosum, subacaule; foliis radicalibus longe petiolatis
triplicato-3-partitis,
laciniis linearibus divaricatis,
caulinis linearibus plerumque indivisibus,
caule superne nudo unifloro.
compositum
On the banks of the Kooskoosky.
M. Lewis.
♃.
July, Aug. v. s.; v. v. cultum.
Not above a span high;
flowers resembling a daisy exceedingly;
they change during their flowering,
from white to a lively pale red.

Coll. No. 1901, Erigeron compositus
• Golden Checklist Flora:
Erigeron divergens;
Erigeron divergens Torr. & A. Gray. Spreading Fleabane.
- Nuttall, Thomas, 1840-1841.
• Glossary:
fastigiate;
Erigeron *divaricatum;
☉
hirsute, stem branching from base;
branchlets one-flowered, rather naked, fastigiate;
radical leaves spathulate, the rest linear, sessile, acute, attenuated below, all entire;
inner pappus of about eight setæ
rays nery numerous, narrow, white.
Hab. In the Rocky Mountains and the plains of Oregon.
About one foot high, at length very much branched,
the leaves an inch or more long, about a line wide.
Pappus double in ray and disk,
the inner of remarkably few rays, very deciduous.

Coll. No. 2092, Erigeron divergens
- Torrey, John, and Asa Gray, 1838-1843.
• Field Notes:
Torrey & Gray, 1841;
26. E. divergens:
somewhat hoary with a minute hirsute pubescence,
diffusely branched from the base;
leaves small, entire, acute;
the radical somewhat spatulate, narrowed into a short petiole;
the cauline scattered, sessile, linear, narrowed at the base;
heads (small) mostly solitary terminating the naked branchlets or pecuncles;
rays very narrow and numerous, twice the length of the hirsute involucre;
inner pappus of few (8-12) very slender and deciduous bristles.
—
Erigeron (Oligotrichum) divaricatum, Nutt. ! In trans. Amer. Phil. Soc. (n. ser.) 7. p. 311, not of Michx.
Rocky Mountains, and plains of the interior of Oregon, Nuttall !
— ①
Stems about a foot high, diffuse, ascending, slender, at length much branched.
Leaves half an inch to an inch long, 1 to 2 lines wide, mucronate-acute.
Heads rather smaller than in E. tenue; the rays (white, Nutt.) nearly similar;
the exterior pappus shorter.
• Golden Checklist Flora:
Erigeron flagellaris;
Erigeron flagellaris A. Gray. Trailing Fleabane.
- Gray, Asa, 1849.
• Field Notes:
Gray, A., 1849;
334. E. flagellare (sp. nov.):
bienne? striguloso-puberulum, pumilum;
caulibus gracillimis e basi ramosis,
floriferis seu primariis simplicibus superne aphyllis monocephalis, sterilibus patentibus flagelliformibus;
foliis spathulatis mucronulatis inferioribus in petiolum attenuatis integris seu radicalibus parce inciso-lobatus,
ramealibus parvulis sublinearibus sessilibus;
ligulis numerosis gracilibus (albis purpureo tinctis) involucrum hirsutum duplo superantibus;
pappo radii et discii conformiduplici,
exteriore coroni-forme-squamellato conspicuo, interiore e setis sub-20 fragilibus.
—
Low, moist places, along Santa Fe Creek; May, June.
(381.)
—
Root slender.
Flowering stems 5 to 7 inches high, very slender, few-leaved below the middle, naked, and pedunculiform above;
the head rather than the preceding species;
the involucre, &c., similar.
Lower leaves one to two inches long, including the slender petiole;
those of the runner-like sterile branches decreasing to 2 or 3 lines in length.
This species should rank next to the foregoing.*
• Plainview Road:
near coll loc;
• Field Notes:
Coll. No. 1116, 28 May 2015;
Coll. No. 1829, 16 May 2018;

Coll. No. 1116, Erigeron flagellaris
Coll. No. 1829, Erigeron flagellaris
- Nuttall, Thomas, 1818.
• Publication Details:
Nuttall, 1818, publication details;
Erigeron pumilus Nutt. Shaggy Fleabane.
- Nuttall, Thomas, 1818.
• Publication Details:
Nuttall, 1818, publication details;
Original Text
Comments
2. *pumilum.
Hirsute;
stems aggregated, 1-flowered, leaves obling-linear, entire and sessile;
flower large and hemispherical, before flowering nutant;
calix very hirsute.
Hab.
On the plains of the Missouri.
Flowering in May.
E. hisutum. Ph. 2. Suppl. p. 742.
but this name has been previously employed for another species.
Obs.
Stems several from the same root, often, indeed, connected at the base,
4 or 5 inches high, 1-flowered.
Leaves 2 inches, more or less, nearly linear, more or less, nearly linear,
attenuated downwards, scarcely 2 lines wide.
Flower naked, (or pedunculate) white, and large as a Daisy;
rays as long as the calix, narrow and numerous.
Pappus double, internal short, about 12-rayed.
- Nuttall, Thomas, 1818.
• Golden Checklist Flora:
Erigeron pumilis;
• Glossary:
nutant;
• Publication Details:
Nuttall, 1818, publication details;
“nutant”, adj., drooping or nodding.
Syngenesia. Superflua. 147
557. ERIGERON. L. (Flea-bane.)
Calix imbricated, subhemispherical, in fruit often reflected.
Rays of the corolla linear, very narrow, and numerous.
Receptacle naked.
Pappus double, exterior minute, interior pilose, of few rays. (12 to 25?)
Herbaceous;
stems simple or branched, flowers solitary, axillary and terminal, subcorymbose or fastigiately paniculate;
radii white, or purplish, rarely destiture of pappus.
Seed very small, rather smooth, oblong, and compressed;
pappus simply pilose, deciduous, not much longer than the seed,
exterior pappus minute and paleaceous.
§ I. Stem simple
SPECIES.
1. E. alpinum.
2. * pumilum.
Hirsute;
stems aggregated, 1-flowered, leaves oblong-linear, entire and sessile;
flower large and hemispherical, before flowering nutant;
calyx very hirsute.
HAB.
On the plains of the Missouri.
Flowering in May.
E. hirsutum.
Ph. 2. Suppl. p. 742.
but this name has been previously employed for another species.
OBS.
Stems several from the same root, often, indeed, connected at the base, 4 or 5 inches high,
1-flowered.
Leaves 2 inches, more or less, nearly linear, attenuated downwards, scarcely 2 lines wide.
Flower naked, (or pedunculate) white, and large as a Daisy;
rays as long as the calix, narrow and numerous.
Pappus double, internal short, about 12-rayed.
…
• Golden Checklist Flora:
Erigeron strigosus;
Erigeron strigosus Muhl. ex Willd. Prairie Fleabane.
- Willdenow, Carl L., 1797-1830.
11. ERIGERON ʃtrigoʃum.
E. foliis lanceolatis utrinque attenuatis ſubdentatis ſtrigoſo-piloſis,
floribus corymboſo-paniculatis. W.
Erigeron ſtrigoſum. Mühlenb. In litt.
Behaartes Berufungskraut. W.
Habitat in Penſylvania. (v. ʃ.)
Caulis erectus ʃtriatus, pilis ʃparʃis albis obʃitus.
Folia alterna lanceolata baʃ et apice attenuata, utrinque oilis copioʃis adpresʃis obʃita,
integerrima, vel medio utrunque ʃerraturis 2. ʃ. 3 inʃtructa.
Flores corymboʃo-paniculati.
Radius copioʃus filiformis albus. W.
- DeCandolle, Augustus Pyramus, 1836.
5. S. strigosa,
caule erecto sparse et patentim piloso,
foliis lanceolatis utrinque stteniatis dentatis strigoso-hispodid, capitulis laxe corymboso-paniculatis,
invol. Glaberrimo, ligulis numerosis filiformibus.
② ? In Pensylvania (Muhl.),
Noveboraco (Torrey!).
Erigeron strigosum Muhl. In Willd. Sp. 3. p. 1956.
Erig. Australe Horn. Ex Spreng.
An Erig. Strigosun Ell. Sketch 2. p. 394 et Doronicum ramosum Walt. Fl. Car. 206 in Carolina cresc. Ad hanc speciem referenda?
(v. s. comm. A cl. Torr.)
- Nesom, Guy L., 2008.
• Golden Checklist Flora:
Erigeron tracyi;
Erigeron tracyi Greene. Running Fleabane.
- Gray, Asa, 1849.
• Publication Details:
Gray, 1849, publication details;
333. E. cinereum (sp. Nov.):
bienne?
Undique molliter cinereo-pilosum:
caule e basi ramoso;
ramis adsurgentibus apice longe nudis monocephalis;
foliis spathulatis vel lineari-oblongis basi attenuatis integerrimis seu radicalibus paucidentatis incisisve;
ligulus numerosissimis gracilibus (albis nunc purpureo tinctis) involucrum hirsutum duplo superantibus;
pappo radii et disci conformi duplici, exteriore coroniformi-squamellato,
interiore e setis sub-20 fragilibus deciduis.
—
Var. a. Is a dwarf, vernal form, only a span high, quite hoary, the primary flowering stems erect and almost scapiform
(no. 374 of the distribution).
Dry, exposed places around Santa Fe; May.
Var. ß has taller and more diffuse stems (10 inches high),
the leaves almost lanceolate, entire, the lower tapering into slender petioles.
Low, sandy banks of the Rio del Norte and of Santa Fe Creek; May to June. (380.)
Var. ? is a larger, coarser, and much more leafy state;
from the valley of Santa Fe Creek, near irrigating ditches; May to July. (385.)
—
The heads are as large as those of Bellis perennis, solitary on peduncles, or the naked summit of the stems,
of from 2 to 4 inches in length.
The species belongs to the first division of the section Phalacroloma, Torr. & Gray, l. c. †
Some forms of this, or of an allied species (possibly E. affine, DC.),
with rather less numerous and white rays, and either entire or incised leaves,
were gathered at Buena Vista and Encantada by Dr. Gregg.
- Gray, Asa, and Charles Wright, 1852.
268. E. divergens, var. cinereum.
E. cinereum, Gray, Pl. Fendl. p. 68.
New Mexico; the locality not recorded.
E. divergens, var. cinereum, Gray, Pl. Wright, l. c.
Moutains, around the copper mines; Oct: a late, much-branched state. (1169)
Hills near El Paso; March, April: the early normal form (1398.)
- Buckley, S. B., 1861.
Northern Texas. May.
Very much branched from the root.
Stems erect, 4-6 inches high;
leaves numerous near the root and upwards on the stems about 3 inches;
the upper stems, two inches below the flowers, naked;
whole plant strigose hirsute, the white hairs not appressed;
flowers about 1/4 of an inch in diameter;
leaves 1/2 - 1 inch long and a little more than a line wide.
- Greene, Edward Lee, 1902a.
• Field Notes:
Greene, E. L., 1902;
Erigeron commixtus.
With the habit of E. flagellaris, smaller, less stoloniferous,
the leaves relativelt broader, some entire,
others with one or more conspicuous lobes at base of the blade,
the whole plant almost hoary with stiffly hirsute pubescence,
this spreading on the leaves and petioles, retrorse on the stems and peduncles:
heads, rays, achenes, etc., much as in E. flagellaris.
Cańon of the Limpia, mountains of western Texas, 26 April 1902, S. M. Tracy and F. S. Earle;
also collected by the present writer, in the same general region,
namely in the mountains near Silver City, New Mexico, 18 May 1880, and distributed for E. flagellaris.
The species last named has a rather obscure, fine closely appressed hairiness.
That of the new one is so extremely different,
that were the plants the size of a Sunflower or Goldenrod,
no botanist would confuse them as one species,
were the pubescence the only character.
I may remark that true E. flagellaris reaches the mountains of even southern New Mexico,
where, however, it occurs only in a more elevated biological zone
- Greene, Edward Lee, 1902a.
- Nesom, Guy L., 2004.
Erigeron Tracyi.
Allied to the last (Erigeron commixtus — Ed.) but dwarf,
only 2 or 3 inches high, densely leafy at base and with no stolons (at least at flowering time):
petioles of the spatulate-lanceolate entire leaves shorter than the blade or obsolete;
the whole herbage silvery-hoary with a fine dense strigulose pubescence,
or this more sparse and spreading on the solitary scapiform peduncle:
involucre hispidulous:
outer pappus very conspicuous though consisting of only shorter and setiform hairs,
the inner of a few very delicate capillary ones.
Davis Mountains, western Texas, Tracy and Earle, 28 April 1902.
- Coulter, John M., and Aven Nelson, 1909.
• Glossary:
cinereous;
33. Erigeron colo-mexicanus A. Nels.
Stems few to several from s slender annual taproot, ascending,
naked pedunculiform above the middle,
very leafy, flowering when very short, the later heads on stems 7-15 cm. long:
leaves mostly linear-spatulate to linear, the basal sometimes oblanceolate and 3-lobed at apex,
all softly cinereous-pubescent:
heads musc as in E. divergens.
(E cinereus Gray, Pl. Fendl. 68. 1848; not E. cinereus H. & A. Comp. Bot. Mag. 2: 50. 1836)
—
Southern Colorado and New Mexico.
- Rydberg, Per Axel, 1910.
Erigeron commixtus Greene, Pittonia 5: 58. 1902
E. cinereus A. Gray, Mem. Am. Acad. 4: 68. 1849. Not E.cinereus H. & A. 1836.
E. colo-mexicanus A. Nels,; Coult. & Nels. New Man. Cent. Rocky Mts. 529. 1909.
Another specific name proposed by Professor Nelson, Erigeron colo-mexicanus,
to replace the untenable E. cinereus A. Gray,
is in my opinion rather distasteful.
Fortunately I do not need to use the name,
as the same species has been described by Dr. Greene under the name E. commixtus.
- Tidestrom, Ivar, 1925.
7. Erigeron nudiflorus Buckley, Proc. Acad. Phil. 1861. 456. 1862.
Erigeron cinereus A. Gray, Mem. Amer. Acad. n. ser. 4: 68. 1849. Not E. cinereus Hook. & Arn. 1836
Erigeron divergens cinereus A. Gray, Pl. Wright. 1: 91. 1852.
E. commixtus Greene, Pittonia 5: 58.1902.
Erigeron colo-mexicanus A. Nels. in Coulter, New Man. Rocky Mount. 529. 1909.
Artemisia, pinyon and yellow pine belts; Utah and Colorado to Mexico.
- Tidestrom, Ivar, and Sister Teresita Kittel, 1941.
9. Erigeron nudiflorus Buckley, Proc. Acad. Phil. 1861. 456. 1862.
E. commixtus Greene, Pittonia 5: 58.1902.
Artemisia, Pinyon and Yellow Pine belts; Utah and Colorado southward to Mexico.
- Cronquist, Arthur, 1947.
- Gray, Asa, and Charles Wright, 1852.
which leads to E. flagellaris. The other half of the couplet:
6. Some or all of the hairs of the stem appressed or closely ascending, or the stem glabrous.
leads to E. divergens where E. tracyi is treated as a variety.
6. Hairs of the stem all spreading.
Key to the varieties of Erigeron divergens
1. Earliest heads on leafy peduncles; plant without long stoloniform branches.
A. var. typicus.
1. Earliest heads on long naked peduncles; plant later producing long leafy stolons or stoloniform branches.
B. var. cinereus.
112B. Erigeron divergens var. cinereus A. Gray, Pl. Wright. 1: 91. 1852.
E. cinereus A. Gray, Mem. Am. Acad. II 4:68. 1849. Not H. & A. 1836.
E. nudiflorus Buckl. Prec. Acad. Phil. 1861: 456. 1862.
E. commixtus Greene, Pittonia 5:58. 1902.
E. tracyi Greene. Pittonia 5: 59. 1902.
E. divergens nudiflorus A. Nels. Man. Bot. Rocky Mts. 529. 1909.
E. colo-mexicanus A. Nels. Loc. Cit.
? E. dicladus Greene. Leafl. 2: 214. 1912.
Earliest heads on long naked peduncles;
plant later producing long loafy stolons or stoliform branches;
basal leaves more often persistent than in var. typicus.
Southern Nevada and Utah, through Arizona and New Mexico to northern Mexico, central Texas, western Oklahome, and southwestern Kansas;
reported from Colorado and western Arkansas.
Type: Fendler 374, near Santa Fe, New Mexico, 1847 (NY).
…
Texas:
Tracy & Earle 279, Limpia Canyon, April 26, 1902 (type of E. commixtus Greene) (MI, NY);
Tracy & Earle 320, Davis Mountains, April 28, 1902 (type of E. tracyi Greene) (MI, NY);
…
Colorado:
Rydberg & Vreeland 5438, 5 miles southwest of La Veta, Huerfano Counto, May 22, 1900 (NY);
…
The variety cinereus has usually been treated as a distinct species under the name E. nudiflorus
Buckl., but there are two many intermediates for it to stand.
There are occasional intermediates with E. modestus,
having the pubescence of the latter, and habit of E. divergens var. cinereus.
These plants are more likely to have conspicuously lobed basal leaves than is typical var. cinereus.
- Harrington, H. D., 1954.
- Harrington, H. D., 1964, 2nd ed..
8B. Erigeron divergens cinereus A. Gray, (var.) Pl. Wright. 1:91. 1852.
E. cinereus A. Gray;
E. nudiflorus Buckl.;
E. commixtus Greene;
E. colo-mexicanus A. Nels.;
E. divergens nudiflorus (Buckl.) A. Nels.
--
Earliest heads on long naked or nearly naked peduncles;
plants producing long leafy stolons later in the season.
---
Dry often sandy soil.
Kansas ro Nevada, south to Texas and Arizona.
Our records scattered over Colorado, mostly in the southern part, at 4000-8000 feet.
- Nesom, Guy L., 1989.
Original Text
Comments
Erigeron tracyi an earlier name for Erigeron colomexicanus
I have been using Erigeron colomexicanus as the name for this species,
but both E. tracyi and E. commixtus were published seven years earlier,
simultaneously (immediate succession in the same paper: Greene 1902).
Plants of the type collections of E. tracyi, as well as those of E. commixtus,
are early season forms
(essentially a basal rosette with a single, subscapiform, monocephalous stem)
that had not yet produced runners characteristic of the species.
Erigeron tracyi Greene, Pittonia 5:59. 1902. Type: U.S.A. Texas. [Jeff Davis Co.:] Davis Mts., 28 Apr 1902,
S. M. Tracy and F. S. Earle 320 (Holotype: US!; Isotypes: GH!, NY!, OC!).
Erigeron commixtus Greene, Pittonia 5:58. 1902. Type: U.S.A. Texas. [Jeff Davis Co.:] Cańon of the Limpia, Mountains of west Texas, 26 Apr 1902,
S. M. Tracy and F. S. Earle 279 (Holotype: US!; isotypes: GH!, NY!, TAES!, TEX!).
Erigeron cinereus A, Gray, Mem. Amer. Acad. Arts n.s., 4 [Pl. Fendler.]; 68. 1849 (not Hook. & Arn. 1836).
Erigeron divergens Torrey & A. Gray var. cinereus (A. Gray) A. Gray, Smithsonian Contr. Knowl. 3, Art. 5 [Pl. Wright.]:91. 1852.
Erigeron colomexicanus A. Nels. [nom. Nov.], Man. Bot. Rocky Mts., 529. 1909. Type: U. S. A. NEW MEXICO. [Santa Fe Co.:] near Santa Fe, 1847,
A. Fendler 374 (Holotype: GH!; isotypes: GH, NY! UC-2 sheets!, US!).
- Flora of North America Editorial Committee, eds., 1993+.
160. Erigeron tracyi Greene, Pittonia. 5: 59. 1902.
Running fleabane
Erigeron cinereus A. Gray 1849, not Hooker & Arnott 1836;
E. colomexicanus A. Nelson;
E. commixtus Greene;
E. divergens Torrey & A. Gray var. cinereus A. Gray
Annuals, biennials, or short-lived perennials, 2.5–8(–12, 18) cm;
usually taprooted, sometimes fibrous-rooted, caudices simple or branched.
Stems first erect (greenish proximally), then producing herbaceous, leafy,
prostrate runners (stoloniform branches, sometimes with rooting plantlets at tips),
densely hirsutulous (hairs spreading-deflexed, of relatively even lengths and orientations),
sparsely minutely glandular.
Leaves mostly basal (persistent in early season);
blades oblanceolate to spatulate (obovate-elliptic laminae), 10–30(–60) × 3–6(–12) mm,
cauline abruptly reduced distally, margins entire, dentate, or lobed, faces densely hirsute, eglandular.
Heads 1(–3 rarely, from midstem or proximal branches).
Involucres 3.5–4.5(–6) × 6–9(–12) mm.
Phyllaries in 3–4 series, sparsely to moderately hirsute, minutely glandular.
Ray florets 60–130; corollas white, often purplish abaxially,
sometimes with an abaxial midstripe, 5–9 mm, laminae not coiling or reflexing.
Disc corollas 2–3 mm (throats indurate and slightly inflated).
Cypselae 0.7–1.3 mm, 2-nerved, faces sparsely strigose;
pappi: outer of setae, inner of 12–16 bristles. 2n = 27.
Flowering Mar–Oct.
Desert scrub, grassy slopes, oak chaparral, pinyon-juniper woodlands, Douglas fir-ponderosa pine;
700–2300(–2400) m; Ariz., Colo., Kans., Nev., N.Mex., Okla., Tex., Utah; Mexico (Baja California, Chihuahua, Coahuila, Durango, Sonora, Zacatecas).
In March through June, plants of Erigeron tracyi produce leaves in a basal rosette usually with a single, erect, monocephalous, stem.
Stoloniform branches are soon formed (often recognized on pressed specimens by the leaves mostly on one side of the branches),
and by the end of the season (August through October), prostrate runners are usually evident, sometimes forming terminal, rooting plantlets.
Erigeron tracyi is similar in habit to E. flagellaris, particularly in the herbaceous stolons or stoloniform branches;
the stem pubescence of E. tracyi is different, the stolons much less commonly produce rooting plantlets at the tips,
and the plants tend to be perennial with woody or lignescent caudices, although they are variable both in habit and duration.
Apparent hybrids with E. modestus and E. flagellaris are occasionally encountered,
and the most common form of E. tracyi is perhaps (speculative) a stabilized, apomictic hybrid between the latter and E. divergens.
All chromosome counts thus far have shown E. tracyi to be triploid and asynaptic.
- Snow, Neil, 2009.
Erigeron tracyi Greene - FNA
[Erigeron cinereus A. Gray 1849 {not Hook. & Arnott 1836} - FNA
[Erigeron colomexicanus A. Nelson] - KTZ, E
[Erigeron colo-mexicanus A. Nelson] - SFE, W&W
[Erigeron commixtus Greene] - FNA
[Erigeron divergens Torr. & A. Gray var. cinereus A. Gray] - HDH, UTF
[Erigeron divergens Torr. & A. Gray var. cinereus (A. Gray) A. Gray] - IMF
[Erigeron modestus A. Gray] - GPF
- Weber, William A., and Ronald C. Wittmann, 2012.
11a. [10] Heads solitary on elongate, mostly leafless stems from the basal leaf clusters;
plants developing long, nonrooting, stolon-like, spreading shoots.
E. tracyi Greene [for Samuel Mills Tracy, 1847-1920].
Dry, gravelly floodplains and meadows, mimicking E. flagellaris but with spreading stem hairs.
Flowering very early in the spring.
The FNA suggests that this is a triploid apomictic hybrid between E. divergens and E. flagellaris.
(E. colomexicanus is a later name.)
- Ackerfield, Jennifer, 2015.
Erigeron tracyi Greene, Running Daisy. [E. colomexicanus A. Nelson].
Annuals, biennials, or short-lived perennials, 1-4 dm.;
leaves oblanceolate to linear, entire or few-toothed, the stems forming stoloniferous branches, 1-3(6) cm. long;
involucre 3-5 mm. hih, hirsute with curved hairs;
disk flowers 2-2.5 mm. long;
ray flowers 4-6 mm. long, white or pink;
pappus double.
Common on the eastern slope in open meadows and on dry slopes, with scattered occurrences on the estern slope, 4000-9000 ft. April-July. E/W
24a Stems with spreading or tangled (pointing in all directions) hairs or mostly glandular … 25
24b. Stems with appressed or ascending hairs (at above the middle), not glandular … 34
• Golden Checklist Flora:
Erigeron vetensis;
Erigeron vetensis Rydb. Early Bluetop Fleabane.
- Rydberg, Per Axel, 1905.
Erigeron vetensis sp. nov.
Densely cespitose-puvinate perennial ; stems 5-8 cm. high,
hirsute, few-leaved ; leaves linear or linear-oblanceolate, hirsute,
2-4 cm. long; heads solitary, about 7 mm, high ; bracts hnear,
acuminate, hirsute as w^ell as slightly glandular-puberulent ; rays
purple, 8-10 mm. long, over i mm. wide; achenes strigose;
pappus more or less double.
In dry places on high mountains of southern Colorado at an
altitude of 2400-3000 m. It is intermediate between E. radicatus
and E. glandulosus, resembling the former most in pubescence,
and the latter in habit.
Colorado: Mountains near Veta Pass, 1900, Rydberg & Vreeland
5427 (type); Veta Mountain and Ojo, 5421, 5422; West
Spanish Peak, 5424.
• Notes on Colorado Flora:
Erigeron subtrinervis;
Erigeron subtrinervis Rydb. ex Porter & Britton “Threenerve Daisy”
• Golden Checklist Flora:
Gaillardia aristata;
Gaillardia aristata Pursh “Blanketflower”
- Pursh, Frederick, 1814.
- Schreber, Johann Christian Daniel van, 1791.
Original Text
Comments
656. GALARDIA. Gen. pl. 1323.
FromPursh's Index Auctorum:
“Gen. pl. — Car. a Linne, Genera Plantarum, Ed. 8vo. curante J. C. D. Schreber.
Vol. 2. Francof. 1789. 1791. 8vo.”
…
3. G. hirsutissima, scabra ; caule ramoso, foliis lineari-lanceolatis
sessilibus, paleis pappi (5—7) ovato-lanceolatis
longissime aristatis.
aristata.
On dry hills on the Rocky-mountains. M. Lewis. ♂.
v. s. in Herb. Lewis.
Flowers orange-coloured.
Planta hirsutissima.
Caulis erectus, teres, ramosus.
Rami uniflori.
Folia caulina alterna, longe-lineari-lanceolata, sessilia, scabra, hirsutissima.
Calyx sub-imbricatus, polyphyllus : foliolis lineari-oblongis,
acutissimis, disco duplo longioribus, margine coloratis.
Corollulae : Radii circiter 12. obcuneatae, 3-fidae :
laciniis oblongis, subaequalibus, obtusiusculis;
Disci tubulosae, 5-fidae : laciniis extus purpurascentibus hirsutissimis.
Receptaculum in ambitu subpaleaceum, centro setosum.
Pappus : paleis 5—7. scariosis, ovatis, longissime-setaceo-aristatis.
• Golden Checklist Flora:
Galinsoga parviflora;
Galinsoga parviflora Cav. “Gallant Soldier”
- Cavanilles, Antonio Jose, 1791.
Original Text
Comments
GALINSOGA PARVIFLORA. tab. 281.
308. GALINSOGA caule herbáceo: foliis oppositis, ovato-lanceolatis, subserratis: pedunculis
elongatis uni, bifloris.
(a) Le réceptacle (loquitur de Amello tenuifolio)
est chargé de paillettes , ainsi que chaque semenee ,
dont l'aigrette n'est pas formée par des
poils: ce caractčre indique un rapport marqué entre
les deux premičres espčces de ce genre, et les Verbesines.
Verbesina biflora. Hort. Reg. Parisiens.
Caulis herbaceus, glaber, bipedalis, ramis oppositis, iterum ramosis.
Folia opposita, ovato-lanceolata, subserrata, trinervia, glabra, nervis exceptis: petioli breves, subconnati.
Flores axillares et terminales, pedunculis elongatis uni aut bifloris.
Calix glaber, hemisphaericus, imbricatus squamulis ovatis dupli serie: harum quinque exteriores carinatae, apice scariosae; interiores vero planiusculae coloratae.
Radii fere semper quinque, aliquando septem, quorum corolla brevis alba:
hermaphroditorum corolla lutea, quemadmodum anthera et stigmata.
Receptaculi paleae apice bi-trifidae, nonnumquam integrae; papi vero brevissimae, sublineares, ciliatae.
Habitat in Peruvia. O Floret Septembri.
Explěc. tab. a Flos integer. b Calix. c Radius auctus. d Flosculus disci auctus.
e Semen. f Idem auctum. g Papi palea. h Receptaculi paleae.
Obs. Hanc plantam vidi in Regio horto Parisiensi anno 1785 nomine Verbesinae biflorae,
ibi enata ex seminibus e Peruvia missis a D. Dombeyo : in patriam redux vidi eam
iterum in Regio horto Matritense, cuius nomen tandem mutatum fuit in debitum Galinsogae.
Obs. I saw it in the Royal Garden of Paris, in the year 1785 given the name of Verbesina biflorae,
arisen from the seeds of Peru, which were sent by D. Dombeyo from there.
I also saw it in my own country in the Royal Garden of Madrid, under a changed name,
that of Galinsoga.
Grindelia Willd.
- Moore, Abigail J., Adriana Bartoli, Roberto D. Tortosa, & Bruce G. Baldwin, 2012.
- Neson, Guy L., 2025.
• Golden Checklist Flora:
Grindelia inornata;
Grindelia inornata Greene “Colorado Gumweed”
- Greene, Edward Lee, 1896.
GRINDELIA INORNATA. Stout and low, the several stems
decumbent, the root perennial: both the short-petioled radical
and broad-based sessile cauline leaves of oval or obovoid
outline and saliently serrate-toothed all around, obtuse at
apex: corymbose-panicled heads large, hemispherical, ray-less:
bracts of the involucre numerous, all squarrose: outer
achenes only somewhat thicker than the thin inner ones, all
truncate at su mmit, only very finely striate: pappus-awns 2,
barbellate below the very acute apex.
Very frequent in southern Colorado, at Cańon City, etc.;
evidently forming a part of Gray's G. squarrosa, var. nuda,
though not at all the G. nuda of Wood. It is most certainly
distinct from G. squarrosa not so much by its discoid heads
as by its perennial root. Not only this but also both the
preceding new species have a persistent tap root surmounted
by a central tuft of radical leaves, around which tuft the
stems of each succeeding year are produced. G. squarrosa,
as I have noted during many seasons, is always strictly biennial,
the stem erect and terminal to the root-axis, the radical
leaves dying as the plant approaches maturity, quite as in
all biennials. Herbarium specimens usually fail to show
these things, hence much confusion in the herbaria and in
books.
- Rydberg, Per Axel, 1906.
- Steyermark, Julian A., 1934a.
- Steyermark, Julian A., 1934b.
- Steyermark, Julian A., 1937.
- Harrington, H. D., 1964, 2nd ed..
- Strother, John L., and Mark A. Wetter, 2006.
- Weber, William A., and Ronald C. Wittmann, 2012.
- Ackerfield, Jennifer, 2022.
- POWO, 2021 - 2026.
- World Flora Online, 2025.
• Golden Checklist Flora:
Grindelia squarrosa;
• Taxa Notes:
Notes on Grindelia, Willdenow, 1807;
Grindelia squarrosa (Pursh) Dunal “Curlycup Gumweed”
• Taxa Notes:
Notes on Grindelia squarrosa;
- Pursh, Frederick, 1814.
Original Text
Comments
637. DONIA. Ait. kew. ed. 2. t. 4. p. 82.
1. D. herbacea ; foliis oblongis amplexicaulibus serratis, squarrosa .
calycinis squamis apice filiformibus revoluto-squarrosis.
squarrosa.
In open prairies, on the banks of the Missouri.
M. Lewis.
♃
Aug. Sept. v. s. in Herb. Lewis. ; v. v. cult.
Flowers yellow, resembling those of Inula.
The whole plant is viscous, and has a strong
resiniferous or balsamic scent.
Caulis erectus, teres, albidus, corymboso-ramosus.
Rami alterni, angulati, apice uni- aut triflori.
Folia alterna, basi dilatata, amplexicauiia, oblonga,
acuta aut obtusa, serrata, glabra,
utrinque squamulis punctiformibus resiniferis obsita.
Flores 1 — 3. in summitate ramulorum, corymbosi, flavi,
ante explicationem copiose resiniferi.
Calyx hemisphaericus, arcte imbricatus :
Squamis apice filiformibus, revolutis et quasi
caput Medusae referentibus.
Flosculi radii lineari-lanceolatae, acutae.
Semina oblongo-obovata.
Pappus: aristis 2 — 4. deciduis.
Receptaculum planum, nudum, favulosum.
It approaches near to Aster glutinosus Cav. ic. 2. p. 53. t. 168.
which is the Donia glutinosa Ait. kew. ed. 2. t. 4. p. 82.
which I distinguish by the following diagnosis :
Donia glutinosa : frutescens ; foliis oblongo-obcuneatis sessilibus serratis,
calycinis squamis linearibus erectis.
- Dunal, Felix, 1819.
Original Text
2. Grindelia squarrosa.
G. caule herbaceo , foliis oblongis amplexicaulibus serratis, involucri squamis apice filiformibus revoluto-squarrosis.
Donia squarrosa. Pursh. fl. bor. am. 2. p. 559*.
—
Sims. bot . mag. t. 1706.
—
Desfont. Tab. ed. 2. p. 269.
Hab. in pratis apricis secůs rivos fluminis Missouri. ( Pursh. ) ♃
Fl. aug. sept.
Caulis erectus teres albidus corymboso-ramosus.
Rami altérai an gulati apice 1-3 flori ex Pursh; omnes 1-flori in speciminibus mihi obviis.
Folia alterna sessilia basi dilatata amplexicaulia oblonga acuta
(interdum obtusa ex Pursh.), serrata , glabra , utrinque squamulis punctiformibus glutiniferis obsita.
Authodia flava ante explicationem copiosč glutinifera erecta corymbosim disposita G. glutinosć magnitudine.
Involucrum hemisphćricum, squamis basi arctč imbricatis, apice
filiformibus patulis révolutisve et ideo capitulum squarrosum constituentibus.
Flosculi Iigulati patentes oblongo-lineares acuti integri.
Semina oblongo-obovata , aristis 2-4 deciduis coronata.
Receptaculum planum nudum favulosum.
Etiamsi herbacea propiůs ad G. glutinosam quam ad alias accedit species.
• Golden Checklist Flora:
Grindelia subalpina;
Grindelia subalpina Greene “Subalpine Gumweed”
- Greene, Edward L., 1898.
Original Text
Grindelia subalpina. Low perennial, the stoutish
corymbose-panicled stems seldom a foot high, only sparingly
leafy except at base; lowest leaves oblanceolate, petiolate,
acute, coarsely and remotely incised, usually scabrous-
puberulent; the few cauline obloug-spatulate, rather remotely
and sharply serrate, glabrous: depressed globose
heads rather large ; bracts of the involucre numerous, with
filiform squarrose-spreading tips, the whole very glutinous:
rays numerous, narrow: bristles of the pappus 2 to 4,
slender, barbellulate.
High plains of southern Wyoming, and at subalpine elevations
on the mountains of northern Colorado. Hitherto
confused with G. squarrosa, which is wholly of the plains,
strictly biennial, branching and very leafy, perfectly glabrous,
and with stouter smooth pappus-bristles.
• Golden Checklist Flora:
Gutierrezia sarothrae;
Gutierrezia sarothrae (Pursh) Britton & Rusby “Broom Snakeweed”
- Pursh, Frederick, 1814.
Original Text
Comments
Sarothrae.
32. S. caule angulato scabro inferne nudo, superne corymboso,
ramis summitate paucifloris, foliis linearibus
stricte-appressis obsolete 3-nervibus utrinque scabris,
axillis nudis, ligulis disco duplo longioribus.
On the plains of the Missouri.
M. Lewis.
♃ Sept.
v. s . in Herb. Lewis.
About a span high; resembling
Hypericum Sarothra in general habit very much.
This and the two preceding have a great affinity to one another ;
but I consider them good and permanent species,
as there are no intermediate varieties, which
might lead to the suspicion that all belong to one genuine species.
The two preceding were Solidago lanceolata and Solidago tenuifolia,
which are now treated as synonyms of Euthamnia graminifolia (L.) Nutt.
and Euthamia caroliniana (L.) Greene ex Porter & Britton, respectively.

Habitat of Coll. No. 1289, Gutierrezia sarothrae
- Lagasca y Segura, Mariano, 1816.
Original Text
Comments
G. N. GUTIERREZIA.
Ord. nat. Corymbiferae. Juss.
Juxta Collumeliam Willdenowě.
372. GUTIÉRREZ. linearifolia : foliis linearibus integerrimis sparsis:
floribus corymbosis.
“Hab. in N. H.” — Habitat in Nova Hispania, i.e., Mexico.

Coll. No. 1289, Gutierrezia sarothrae
- Nuttall, Thomas, 1818.
- Torrey, John, and Asa Gray, 1838-1843.
- Britton, Nathaniel Lord, and H. H. Rusby, 1887.
Original Text
Comments
55. Gutierrezia Sarothrae (Pursh.).
(Solidago Sarothrae, Pursh. Flor. Amer. Sept., ii., 540, 1814 :
Bachyris Euthamniae, Nutt., Genera, ii. 163, 1818 :
G. Euthamiae, T. & G.)
- Lane, Meredith A., 1982.
• Golden Checklist Flora:
Helianthus annuus;
Helianthus annuus L. “Common Sunflower”
- Linne´, Carl von, 1753.
Original Text
POLYGAMIA FRUSTRANEA
HELIANTHUS.
annuus.
1. HELIANTHUS foliis omnibus cordatis : nervis pone bafin unitis : extrorsum denudatis.
Helianthus radice annua. Vir. cliff. 88. Hort. cliff. 419. Hort. upf. 268. Roy. lugdb. 180.
Helenium indicum maximum. Bauh. pin. 276.
Herba maxima. Dod. pempt. 264.
Chryfis. Reneal. fpec. 84. t. 83.
Habitat in Peru, Mexico. (.)
• Golden Checklist Flora:
Helianthus nuttallii;
Helianthus nuttallii Torr. & A. Gray “Nuttall's Sunflower”
- Nuttall, Thomas, 1840-1841.
- Torrey, John, and Asa Gray, 1838-1843.
Original Text
* * * * *
Perennial : heads middle-sized : rays 8-24 : involucre irregularly imbricated ;
the scales loose, or with squarrose-spreading often foliaceous summits,
as long as the yellow disk (achenia glabrous).
—
Corona-solis.
†
Leaves commonly alternate or scattered,
the lower often opposite, feather-verined,
sometimes obscurely triplinerved.
16. H. Nuttallii :
stem smooth ; leaves alternate, the lower opposite,
narrowly lanceolate-linear, acute, mostly entire, scarcely petioled,
both sides scabrous ;
scales of the involucre lanceolate-subulate, hirsute-ciliate towards the base;
pappus of 2 linear-lanceolate chaffy awns or scales.
—
H. californicus, Nutt. ! in herb. acad. Philad. &c., not of DC.
Plains of Lewis River, Nuttall ! —
Stem apparently strict and simple.
Leaves 4-6 inches long, 3-5 lines wide, feather veined,
obscurely triplinerved near the base, somewhat cinereous beneath ;
the lower remotely and slightly serrate.
Heads nearly as large as in H. giganteus.
Involucre clothed with whitish hairs, or often smoothish.
The disk-corolla is 5-nerved,
of with intermediate nerves corresponding with the axis of 2 only of the laciniae,
but not reaching the apex.
• Golden Checklist Flora:
Helianthus pauciflorus ssp. subrhomboideus;
Helianthus pauciflorus ssp. subrhomboideus (Rydb.) O. Spring & E. Schilling “Stiff Sunflower”
- Nuttall, Thomas, 1818.
589. HELIANTHUS. L. (Sunflower.)
Calix imbricated, subsquarrose, foliaceous.
Receptacle paleaceous, flat.
Pappus paleaceous, 2-leaved, caducous.
…
9. * pauciflorus.
Leaves opposite, linear-lanceolate, acuminate, serrate, nearly smooth;
stem naked, trichotomous, few-flowered;
calix closely imnricated; leaflets ovate.
HAB.
In Lower Louisiana.
—
Leaves sometimes ternately verticillate, very long, paler beneath and somewhat pubescent;
ray and disk nearly the same colour.
4 or 5 feet high.
- Cassini, Alexandre Henri Gabriel, 1818.
- Kartesz, John T., and Kancheepuram N. Gandhi, 1990a.
Harpalium rigidum, H. Cass.
Tige herbacée, haute d'environ cinq
pieds, dressée, rameuse, cyliudrique, garnie de poils roides. Feuilles
opposées, presque sessiles, lancéolées, pas sensiblement dentées, d'une
substance ferme et roide, d'un vert glauque ou cendré, munies sur
les deux faces de poils courts et roides. Calalhides grandes, solitaires
au sommet des rameaux nus et pédonculiformes ; fleurs jaunes. (Cultivé
au Jardin du Roi.)
Harpalium rigidum, H. Cass.
Stem herbaceous, about five high
feet, erect, branched, cylindrical, furnished with stiff hairs. Leaves
opposite, almost sessile, lanceolate, not noticeably toothed, of a
firm and stiff substance, of a glaucous or ashen green, provided on
both sides of short, straight hair. Calalhids large, solitary
at the top of bare and stalked branches; yellow flowers. (Cultivated
in the King's Garden.)
- Kartesz, John T., and Kancheepuram N. Gandhi, 1990a.
- Sims, John, M.D., 1821.
Helianthus diffusus. Missouri Sun-Flower.
********************
Class and Order.
Syngenesia Polygamia Frustranea.
Generic Character.
Recept. paleaceum, planum. Pappus diphyllus.
Cal. imbricatus, subsquarrosus.
Specific Character.
Helianthus diffusus ; caule hispido divaricato, foliis ovatis
rigidis scabris serrulatis alternis oppositisque, pedunculis
longissimis unifloris.
Descr. Stem angular, purple, hispid :
branches distant,
rambling, bearing one terminal flower on a very long peduncle.
The whole plant is clothed with stiff, stinging hairs,
scarcely exceeds two feet in height, but spreads wide.
Leaves both opposite and alternate, oblong-ovate, rigid, very rough.
Calyx imbricated, scales ovate, in four series, smooth, with villous edges.
Flowers large : rays about twenty, oblong-ovate,
plicate, three-toothed, of a very full yellow colour.
Floscules of the disk yellow: tube filiform pedicle-like :
border cylindrical.
Anthers dark purple.
Stigmas revolute, golden yellow.
Chaff of the receptacle linear-lanceolate, concave, green.
Germens three-cornered, crowned with a two-leaved, awl-shaped pappus.

Sims (1818) Plate 2020 of Helianthus diffusus.
- Nuttall, Thomas, 1834b.
- Nuttall, Thomas, 1840-1841.
• Taxa Notes:
Notes on Helianthus pauciflorus ssp. subrhomboideus, Nuttall, 1841;
- Cuvier, Frederic, 1821.
Harpalion roide ; Harpalium rigidum, H. Cass., Bull. Des Sc.,
septembre 1818. La tige est herbacee, haute d'environ cinq
pieds, dressee, rameuse, cyiindrique, garnie de poils roides.
Les feuilles sont opposees, presque sessiles, lanceolees, pas
sensiblement dentees, d'une substance ferme et roide, d'un
vert glauque ou cendre, rnunies sur les deux faces de poils
courts et roides.Les calathides sont grandes, solitaires au
sommet des rameaux nus et pedonculiformes; les fleurs sont jaunes.
Cette plante est cultivee au Jardin du Roi , ou nous avons observe,
sur des individus vivans , les caracteres generiques et
specifiques que nous venons de decrire. M. Desfontaines pense
que c'est l'helianthus diffusus, plante vivace, de l'Amerique
septentrionale, decrite dans le Botanical Magazine. (H.Cass.)
Stiff Harpalion; Harpalium rigidum, H. Cass., Bull. Of Scs,
September 1818. The stem is herbaceous, about five
feet, erect, branched, cylindrical, furnished with stiff hairs.
The leaves are opposite, almost sessile, lanceolate, not
noticeably toothed, of a firm and stiff substance, of a
glaucous green or ash, with hairs on both sides
short and stiff.
Calathids are large, solitary in
tips of bare and stalked branches; the flowers are yellow.
This plant is grown in the Jardin du Roi, where we observed,
on living individuals, the generic characters and
specific that we have just described. Mr. Desfontaines thinks
that it is the Helianthus diffusus, perennial plant, from northern America,
described in the Botanical Magazine. (H.Cass.)
- Desfontaines, Rene Louiche, 1829, 3rd edition.
Desfontaines make no mention of Nuttall's Gen. Am. or his H. pauciflorus.
Helianthus.
...
{
rigidus Am. S. ♃
diffusus? Bot Mag.
Harpalium rigidum. H. Cas. Dict. V. 20 p. 3000.
- Nuttall, Thomas, 1840-1841.
• Taxa Notes:
Notes on Helianthus pauciflorus ssp. subrhomboideus, Nuttall in Arkansas;
Helianthus * crassifolius; ♃, stem simple, herbaceous, subscabrous; leaves nearly
all opposite, lanceolate, acuminate at either end, serrate, above smooth, beneath
very scabrous, and, as it were, shagreened; stem naked, one to three-flowered,
with a few small, linear leaves; involucrum closely imbricated, the scales ovate,
slenderly and finely ciliate; achenium subquadrangular, with sometimes four
scales, the two central ones smaller.
Hab. Plains of Arkansa. About two feet high, the lower part of the stem very leafy, above
nearly naked from the sudden diminution in the size of the leaves, the plant here and there scattered
with drops of resin. Leaves half a foot long, or more, about an inch wide, very thiick and
coriaceous. Rays about twenty, bidentate. Achenium pubescent above; with small scales from
the inner angles; discal florets as usual, enlarged and pubescent towards the base.
Allied to H. pauciflorus.
- Torrey, John, and Asa Gray, 1841.
11. H. rigidus (Desf.) : stem simple or sparingly branched, rough; cauline
leaves opposite, very thick and rigid, lanceolate or oblong-lanceolate,
acute or acuminate at each end, subsessile, obscurely serrate or entire, somewhat
triplinerved, extremely hispid-scabrous on both sides ; the uppermost
sometimes alternate; the radical oval, obtuse, strongly triplinerved, petioled ;
scales of the involucre ovate, mostly obtuse, finely ciliate, numerous, closely
imbricated; chaff of the receptacle obtuse; achenia somewhat hairy; pappus
of 2 concave lanceolate scales, rather than awns, and frequently with
one or several intermediate small scales.
—
Desf. Cat. Hort. Par. Ed. 3. p. 184.
H. atrorubens, Michx.! Herb, in part;
Hort. Berol. ! 1839 ;
Bot. mag. T. 2668 ;
Hook. ! Compan. To bot. mag. 1. p. 98 (var. foliis acutioribus).
H. scaberrimus. Ell. ! Sk. 2. p. 423 ;
DC. I. c. p. 588.
H. crassifolius, Nutt. ! In trans. Amer. Phil. Soc. I. c.
Harpalium rigidum, Cass, in dict. Sci. nat. 20. p. 300 ;
DC. ! Prodr. 5. p. 583.
…
‡ Obscure or little-known species.
32. H. paucifiorus (Nutt.) : leaves opposite, linear-lanceolate, acuminate,
serrate, nearly smooth ; stem naked, trichotomous, few-flowered ; scales of
the involucre closely imbricated, ovate. Nutt. Gen. 2. p. 177.
- Porter, Thomas C., and John M. Coulter, 1874.
Helianthus rigidus, Desf.
—
Hall & Harbour ; Dr. Smith.
Cańon City, Brandegee.
Platte River, Coulter.
- Gray, Asa, 1884.
“§ 2. Perennials : receptacle convex, or in some at length low-conical :
lower leaves aalmost opposite. ”
H. rigidus, Desf. A foot or two (rarely 6 to 8 feet) high, rigid, sparingly branched : leaves
very firm-coriaceous and thick, both sides hispidulous-scabrous, shagreen-like, entire or
serrate, lightly triplinerved but indistinctly and sparingly veined ; lower oblong and ovate-
lanceolate, attenuate at base into short winged petioles ; upper mostly lanceolate : heads
comparatively large, showy (disk three-fourths inch high) : involucre pluriserially imbricated ;
its bracts mainly ovate, obtuse or acutish, rigid, appressed, densely and minutely ciliate:
rays numerous, generally inch and a half long : akenes oblong-obovate, 3 lines long :
pappus of two large ovate-lanceolate paleae, and sometimes two or four rather stout
intermediate paleae ! more commonly none.
—
Cat. Hort. Par. ed. 3, 184;
Torr. & Gray, Fl. ii. 322.
H. atrorubens, Michx. l. c, in part ;
Bot. Reg. t. 508 ;
Bot. Mag. t. 2668 ;
DC. Prodr. v. 586.
H. diffusus, Sims, Bot. Mag. t. 2020.
H. Missuricus, Spreng. Syst. iii. 618, name in place of diffusus.
H. scaberrimus, Ell. Sk. ii. 423.
H. Missouriensis (Schweinitz)
& H. crassifolius, Nutt. Trans. Am. Phil. Soc. l. c.
Harpalium rigidum, Cass. Dict. Sci. Nat. xx. 200 ;
DC. Prodr. v. 583, founded on the form with intermediate paleae to the pappus.
Plains and prairies, Saskatchewan and Michigan to W. Georgia, Texas, and eastern part of Colorado.
Sometimes the disk-corollas are at first yellow !
- Coulter, John Merle, 1885.
4. H. rigidus, Desf. A foot or two (rarely 6 to 8 feet) high, rigid,
sparingly branched : leaves very firm-coriaceous and thick, both sides
hispidulous-scabrous, shagreen-like, entire or serrate ; lower oblong and ovate-lanceolate,
attenuate at base into short winged petioles ; upper mostly lanceolate : heads
comparatively large, showy; disk ¾inch high, dark purple or brownish:
involucre pluriserially imbricated ; its bracts mainly ovate, obtuse or acutish, rigid,
appressed, densely and minutely ciliate. Plains and prairies from Michigan
to Texas and west to E. Colorado.
- Rydberg, Per Axel, 1900a.
* Helianthus subrhomboideus.
Perennial ; stem 3-6 dm. high, terete, striate, tinged with red,
sparingly hirsute, simple ; leaves opposite, firm, very scabrous, triple-veined,
generally not strongly serrate, the basal broadly ovate
or obovate-spatulate ; stem-leaves rhomboid-ovate or rhomboid-lanceolate,
short-petioled, 5-10 cm. long, the uppermost diminutive, lanceolate ;
heads mostly solitary, sometimes 2 or 3, 1-1.2 cm. high and
1.5-2 cm. in diameter; bracts in 4—5 rows, oblong, acutish, densely
white-ciliate on the margins ; disk dark brown or purplish ; rays
about 1.5 cm. long.
Nearest related to H. scaberrimus, but differs in the broader and
shorter leaves, the less acute bracts, the smaller heads, and generally
longer peduncles. It grows on prairies, up to an altitude of about
1000 m.
Montana: Sand Coulee, 1885, R. S. Williams, 249;
Columbia Falls, Mrs. Kennedy, 6 ;
Judith Mts., 1882, R. W. Springer, XXXV.
Nebraska: Keya Paha River, 1893, Clements, 2866 ;
Long Pine, 1890, G. D. Sweezey, 70;
Whitman, 1893, Rydberg, 1627 (type).
Dakota: Mouth of Big Sioux River, 1853, Hayden Survey;
Upper Missouri, Nicollet;
Custer, 1892, Rydberg, 805.
Assiniboia: Cypress Hills, 1880, John Macoum ;
Souris Plains, 1883, J. M. Macoiun.
Saskatchewan: 1857-8, E. Bourgeau.
- Rydberg, Per Axel, 1906.
5. Helianthus subrhomboideus Rydb. (H. rigidus A. Gray, in part) On
plains from Man., Sask. And Mont. To Colo. And Neb. — Alt. 4000-8000 ft. —
La Pagosa ; mountains, Larimer Co. ; near Pagosa Peak ; Cheyenne Mountain;
Bosworth's ranch, Stove Prairie; Boulder; canon west of Palmer Lake;
Horsetooth Mountain.
- Coulter, John M., and Aven Nelson, 1909.
4. Helianthus scaberrimus Ell. Bot. S. C. & Ga. 2: 423. 1824. Stem 3-12
dm. high, rigid, sparingly branched: leaves very firm-coriaceous and thick,
both sides hispidulous-scabrous, shagreen-like, entire or serrate; the lower ovate
and ovate-lanceolate, attenuate at base into short- winged petioles; the upper
mostly lanceolate: heads comparatively large, showy: disk 15-18 mm. high,
dark purple or brownish ; involucre pluriserially imbricated ; the bracts mainly
ovate, obtuse or acutish, rigid, appressed, densely and minutely ciliate.
H. rigidus. (H. subrhomboideus Rydb. Mem. N. Y. Bot. Gard. 1: 419. 1900.)
Plains and prairies; Montana to Colorado and Georgia.
- Kartesz, John T., and Kancheepuram N. Gandhi, 1990a.
Helianthus.
Helianthus rigidus (Cass.) Desf. (Cat. Pl., ed. 3, 184. 1829) was based
on Harpalium rigidum Cass. (Bull. Sci. Soc. Philom. Paris 141. Sep 1818).
For Helianthus rigidus ssp. rigidus, Heiser (Mem. Torrey Bot. Club 22[3]:131-138. 1969)
cited several synonyms,
including Helianthus diffusus Sims (Bot. Mag. 45:2020. Oct 1818)
and Helianthus paucifiorus Nutt. (Gen. 2:177. 3 Apr 1818).
Cleariy, Helianthus paucifiorus has priority over Harpalium ngidum and
Helianthus diffusus; hence, the latter two cannot serve as basionyms. For the
North American flora, we accept Helianthus paucifiorus Nutt. As the correct
name.
Helianthus.
The new combination Helianthus pauciflorus Nutt. ssp. subrhomboideus
(Rydb.) Spring & E. Schilling (Biochem. Syst. Ecol. 18(1):22. Mar 1990)
predates H. pauciflorus ssp. subrhomboideus (Rydb.) Kartesz & Gandhi
(Phytologia 68(6):423. Jun 1990) and renders the latter to be superfluous. The
March issue of Biochemical Systematics & Ecology was received in May 1990
at UNC-Botany Library, and by this time, our article in Phytologia was
already in press. Nevertheless, we regret the oversight and correct the subspecies
authorship to: (Rydb.) Spring & E. Schilling.
- Spring, Otmas, and Edward E. Schilling, 1990.
H. pauciflorus ssp. subrhomboideus.
The following nomenclatural change is necessitated by the decision [6] that H. rigidus
is not the correct epithet for this taxon:
Helianthus pauciflorus Nutt. ssp. subrhomboideus (Rydb.) Spring and E. Schilling, comb. nov.
Basionym: Helianthus subrhomboideus Rydb. (1900) Mem. N. Y. Bot. Gard. 1, 419.
Synonym: Helianthus rigidus ssp. subrhomboideus (Rydb.) Heiser (1969) Mem. Torrey Bot. Club 22, 136.
Oklahoma, Kay Co., Taylor 28348;
Indiana, Heiser 439 (IU); Iowa, Ames Co., Stolzfus 7259 (IU).
- Nuttall, Thomas, 1840-1841.
• Golden Checklist Flora:
Helianthus pumilus;
• Publication Details:
Nuttall, 1841, publication details;
Helianthus pumilus Nutt. “Little Sunflower”
There are no online data records of the Nuttall collection of his Little Sunflower.
Original Text
Helianthus * pumilus;
♃
hirsutely pilose and scabrous;
leaves ovate-lanceolate, opposite, attenuated below,
subpetiolate, nearly entire, and three-nerved,
upper leaves lanceolate, alternate;
involucrum hoary, hispid;
sepals imbricated, lanceolate, acute,
as well as the receptacular paleae;
achenia smooth.
Hab.
Rocky Mountains and plains of the Platte.
A low, perennial, simple stemmed species,
about a foot high,
leaves two to three inches long, about an inch wide.
Capituli about three to five (apparently) sessile.
Rays about sixteen, longer than the disk,
paleae somewhat obtuse,
hirsute at the summit,
scales of the achenium rather large and wide.
• Golden Checklist Flora:
Heliomeris multiflora;
Heliomeris multiflora Nutt. “Showy Golden Eye”
- Nuttall, Thomas, 1848b.
Original Text
* HELIOMERIS. †
Capitulum many-flowered, heterogamous;
rays ligulate, in a single series, neuter;
discal florets tubular, hermaphrodite.
Involucrum irregularly imbricated and leafy,
in about two series, and rather spreading.
Receptacle conic, the palea embracing the florets,
lanceolate and acute.
Corolla, rays ligulate, (10-12,)
those of the disk tubular, the tube short, throat wide and cylindric,
border five-toothed.
Stigmata with obling tips.
Achenia laterally compressed,
somewhat tetragonous, smooth, and without any pappus.
A perennial tall herb, exactly resembling an Helianthus,
with narrow, entire, somewhat scabrous leaves, the lower ones opposite ;
flowers yellow, terminal, numerous.
H. *multiflorus.
A stoutish perennial, three or four feet high, much branched ;
stems terete and striated ;
leaves narrow, lanceolate, entire, nearly sessile,
above somewhat scabrous, opposite below, on the branchlets alternate ;
flowers terminal, numerous and showy, bright yellow,
with to to twelve rays,
sometimes with deeper discoloured blotches towards the base or lower half,
entire and oblong ;
leaflets of the calyx oblong-lanceolate, somewhat spreading,
in nearly a simple series ;
achenia black and smooth.
Scarcely distinguishable from Helianthus,
except by a short conic receptacle,
and naked seeds.
Hab.
Mountains of Upper California, (Dr. Gambel,)
and was also collected in the Rocky Mountains by Mr. Gordon.
‡ In allusion to its close affinity to Helianthus.
- Harrington, H. D., 1964, 2nd ed..
- Nuttall, Thomas, 1840-1841.
• Golden Checklist Flora:
Heterotheca foliosa;
Heterotheca foliosa (Nutt.) Shinners. “Hairy False Goldenaster”

Coll. No. 1258, Heterotheca foliosa
Inflorescence of Coll. No. 2177, Heterotheca foliosa
- Nuttall, Thomas, 1840-1841.
• Publication Details:
Nuttall, 1841, publication details;
Original Text
Comments
Chrysopsis * foliosa; ♃,
sericeously villous, and more or less canescent,
the margin and lower surface of the leaves scabrous;
flowers fastigiate, corymbose;
leaves entire, oblong or oblong-ovate, subamplexicaule, crowded,
acute, ciliate below;
scales of the involucrum linear, acute, villous;
achenium silky;
pappus scarcely scabrous, outer pappus slender, dimidiate.
Hab.
In the Rocky Mountain plains, near the banks of the Platte.
Flowering in August.
About a foot high, sending up many hairy stems from the same root.
Nearly allied to C. villosa,
but far more pubescent and hoary, with the leaves widest at the base.
In some specimens quite hoary, the hairs feel as soft as silk,
but on removing this clothing,
the under surface is covered with numerous scabrous elevations.
A very showy species.
• Golden Checklist Flora:
Heterotheca villosa;
Heterotheca villosa (Pursh) Shinners “Hairy False Goldenaster”
- Nuttall, Thomas, 1813.
Original Text
Comments
80*Sideranthus integrifolius. ‡ M.
‡ = Perennial
M. = from the Missourie.
- Pursh, Frederick, 1814.
Original Text
650. AMELLUS. Gen. pi. 131 5.
villosus.
1. A. villosissimus ; foliis sessilibus oblongis acuminatis
integerrimis, floribus axillaribus brevi-petiolatis, radiis
integris, paleis setaceis.
On the Missouri. ♃. v. s. Rays yellow.
- Shinners, Lloyd Herbert, 1951.
• Golden Checklist Flora:
Heterotheca villosa minor;
Heterotheca villosa (Pursh) Shinners var. minor (Hook.) Semple “Hairy False Goldenaster”
• Notes on Colorado Flora:
Hieracium albiflorum;
Hieracium albiflorum Hook. “White Hawkweed”
- Hooker, Sir William Jackson, 1829-1840.
4. H. albiflorum ;
caule folioso inferne molliter deflexo-piloso, foliis lato-lanceolatis
integerrimis obtusis in petiolum attenuatis longe albo-pilosis, caulinis remotiusculis sensim
minoribus sessilibus, panicula gracili multiflora glabra, pedicellis divaricatis, involucris pilis
longis albidis sparsis, “floribus albis."
Hab. Alpine woods in the Rocky Mountains, north of the Smoking River, in lat. 56°. Drummond.
—
This has somewhat the habit of H. Gronovii, but it is very different in other particulars.
The leaves are clothed with long, shaggy, very soft white hairs, which are copious, and remarkably deflexed on the
petioles and base of the stem; while the rest of the stem and the panicle are glabrous, and the involucre has
only a few long, scattered, patent hairs.
• Notes on Colorado Flora:
Hieracium fendleri;
Hieracium fendleri Sch. Bip. “Yellow Hawkweed”
- Gray, Asa, 1849.
454. Crepis ambigua (sp. nov. non Balb.):
caule scapiformi e radice perenni simplici basi folioso apice racemoso 5—8-cephalis
foliisque obovati-oblongis integerrimis setis
patentibus sparsis hispidis ceterum glaberrimis;
capitulis majusculis longiuscule pedunculatis circa 30-floris ;
involucro cylindrico hirto, squamis linearibus, extimis duplo brevioribus appressis ;
ligulis (flavis ?) genitalia haud superantibus ;
acheniis fusiformibus superne sensim angustatis multicostatis glaberrimis pappum albidum longitudine equantibus.
—
Level, grassy places along Santa Fé Creek; June to Aug.
— A foot high, from a thick perennial root, which yields a white juice;
the purplish stem leafy only near the base,
above nearly smooth and bearing one or two linear bracts.
Leaves 2 or 3 inches long, sessile ;
the radical with a tapering base, hispid with slender bristles above, purplish underneath,
resembling those of Hieracium Gronovii, β. subnudum.
Heads two thirds of an inch in length.
Ligules very short, little longer than the pappus, usually shorter than the styles.
Mature achenia blackish, a third of an inch long, attenuated upwards, but not truly rostrate.
Pappus copious, dull white, of bristles nearly as stiff as those of Hieracium, but occupying much more than one series;
—
on which account I refer the plant to Crepis (although the pappus is not bright white) rather than to Hieracium.
- Schultz-Bipontinus, C. H., 1861.
Subgen. Chionoracium SzBip. Pappus niveus.
…
B. Crepidicarpum SzBip. Achaenia elongata, fusiformia.
a. Scapiformes, pappus subniveus, subflavescens.
Hieracium Fendleri Sz Bip. = Crepis ambigua A. Gray! in pl. Fendl. p. 114.
- Porter, Thomas C., and John M. Coulter, 1874.
HIERACIUM FENDLERI, Schultz. (Crepis ambigua, Gr. Pl. Fendl., p. 114.)
Stem scape-form and simple from a perennial root, or with 1 or 2
elongated branches from the base or near it, each bearing 5-8 heads ;
leaves obovate-oblong, very entire, beset with scattered, spreading,
hispid bristles, otherwise very smooth, 2' -3' long, sessile, the radical
with a tapering base, somewhat purplish underneath ; heads 9" long, on
rather long peduncles; florets about 30; involucre cylindrical, hairy,
scales linear, exterior ones twice shorter, appressed ; ligules very short,
a little longer than the pappus, usually a little shorter than the styles;
achenia fusiform, narrowed upwards, but not truly rostrate, 4" in length,
many-ribbed, very smooth, as long as the copious, dull-white pappus.
Mace's Hole, Fremont County, Brandegee. Parry.
Wet Mountain Valley, Redfield.
- Fehrer, Judith, et al., 2022.
• Notes on Colorado Flora:
Hieracium triste;
Hieracium triste Willd. ex Spreng. “Slender Hawkweed”
- Sprengel, Curt Polycarp Joachim, 1826.
trifte W. herb.
29. H. foliis erectis longe petiolatis oblongis fubintegerrimis glabris,
caule fuperne anthodiisque hirtis nigrefcentibus.
Inf. Aleut.
triste, Willdenow's herbarium
29. Hawkweed with erect, long-stalked, oblong, nearly entire, smooth leaves;
the upper part of the stem and the flower-heads are hairy and blackish.
South of the Aleutian Islands.
Hymenothrix A. Gray
- Gray, Asa, 1849.
HYMENOTHRIX, Nov. Gen.
Capitulum multiflorum, heterogamum ;
fl. radii 8— 10 ligulatis foemineis, disci tubulosis.
Involucrum turbinatum, disco brevius, e squamis circiter 10 lanceolato-oblongis, subbiseriatis, equalibus, appressis.
Receptaculum parvum, planum, nudum, papillosum.
Ligulae spathulato-oblongae, apice 3-dentate, discum haud superantes.
Corollae fl. hermaph. elongate, tubo gracili minutim glanduligero, fauce infundibuliformi 5-loba
lobisque margine vix incrassatis glabris.
Antherae ecaudate, exserte, appendicula ovata superate.
Styli rami fl, hermaph. lineares, cono brevi superati.
Achenia lineari-cuneata, sub 4—5-gona, glabriuscula.
Pappus simplex, radii et disci conformis, corolla paulo brevior, e setis 12—13 equalibus sursum
barbellulatis infra medium membranula hyalina lanceolata integerrima utrinque limbatis.
&mdas;
Herba annua ? amara, subglabra ;
caule erecto corymboso ; foliis alternis petiolatis biternatisectis, segmentis integris vel trifidis
lobisque linearibus; capitulis corymbosis ;
floribus radii et disci flavis.
H. Wistizeni.
—
Grassy places, Ojo de Gallejo, between El Paso del Norte and Chihuahua, August, Dr. Wislizenus.
—
Stem two feet high, rigid, loosely and amply corymbose at the summit;
the branches, leaves, involucre, &c., minutely appressed-puberulent.
Lower leaves triternately, the upper biternately, dissected ;
those of the branches 3—5-parted, or the uppermost simple;
the segments a line or less in width, half an inch or more in length.
Heads 4 lines long, on slender and minutely bracteate peduncles;
the fully developed flowers nearly twice the length of the herbaceous and appressed involucre.
Disk-corollas 3 lines in length, fully as long as the spreading rays;
the sinuses between the ovate lobes slightly unequal.
Branches of the style narrowly linear, semicylindrical or more flattened, glabrous,
or minutely puberulent on the back, the prominent stigmatic lines extending to the base of the short,
minutely bearded cone with which they are tipped.
Pappus longer than the achenium, formed of narrow and diaphanous paleae with a very strong midrib
which is gradually excurrent into a prolonged, barbellate-denticulate awn.
&mdash.
I know of no genus of Euhelenieae with which this plant can be immediately compared,
except Chaetymenia, Hook. & Arn. (in Bot. Beech., which the authors refer to Tagetineae),
Burrielia, DC., and Oxypappus, Benth.
In habit, foliage, &c., it is very unlike either of these: the pappus is much as in the former;
but the style differs widely from the description and figures of this and the last-named genus.
• Golden Checklist Flora:
Hymenothrix dissecta;
Hymenothrix dissecta (A. Gray) B. G. Baldwin “Ragleaf Bahia”
- Gray, Asa, 1849.
† 425. Amauria ? dissecta (sp. nov.) : hierbacea, puberula ; caule adscendente apice
corymbosi-polycephalo ; foliis alternis petiolatis biternatisectis segmentis cuneiformibus
vel sublincaribus saepius 2-3-fidis, summis parvis ; pedunculis glandulosis ; involucri
squamis oblongo-lanceolatis subtriseriatis, intimis subscariosis ; receptaculo convexo ;
ligulis circiter 16; styli ramis fl. disci cono brevissimo truncati-capitatis ; acheniis ad
angulos laevibus.
—
A few miles east of Mora River ; Aug. (537.) Also gathered in
Fremont's third expedition, probably towards the head-waters of the Arkansas.
—
Stem 12 or 15 inches high, apparently from a perennial root. Leaves about an inch in diameter,
cut into narrow divisions. Peduncles clothed both with viscous and capitate-glandular hairs.
Involucre herbaceous, more or less viscous, a third of an inch in diameter.
Receptacle entirely destitute of chaff. Flowers all yellow: rays linear-oblong, 2-3-toothed ;
the tube very glandular. Disk-corollas with the slender tube extremely
glandular, the expanded 5-cleft limb slightly so. Branches of the style short, flattish-semiterete,
capitate with a very short and flattish obtuse cone. Achenia cuneate-linear,
slender, compressed-quadrangular, smooth ; the ovary sprinkled with sparse and minute
hairs. Pappus none.
—
The specimen of Fendler has not matured fruit ; and the stamens
are abortive in all the disk-flowers. The specimen from Fremont's collection,
communicated by Dr. Torrey, is very imperfect, but has ripe achenia. From the character
of the Californian genus Amauria, Benth. in Bot. Voy. Sulph. p. 31, this plant differs very
essentially in the styles, and in the convex receptacle. But I am unwilling to constitute
it a distinct genus upon the present imperfect materials.*
- Britton, Nathaniel Lord, 1889.
Original Text
Comments
Bahia dissecta (Gray). Amauria (?) dissecta, Gray. Mem. Amer.
Acad. iv. 104 (1849); Villanova chrysanthemoides, Gray,
Smithsonian Contr. v. 96 (1853); Bahia chrysanthemoides,
Gray, Proc. Amer. Acad. xix. 28 (1883). Copper Cańon.
A form with involucral scales not acuminate (183).
It is unclear which Copper Cańon this might be.
There is no Copper Cańon contained within the current named San Francisco Mountains or the Mongollon Rim.
The closest might be Copper Canyon along US Interstate 17 near Camp Verde.
- Rydberg, Per Axel, 1915.
Original Text
Comments
28. AMAURIOPSIS Rydberg, gen. nov.
Glandular-pubescent annuals. Leaves alternate, twice or thrice ternately divided.
Heads in leafy corymbs, radiate. Involucre hemispheric; bracts 16-20, herbaceous, oblanceolate
in about 3 series. Receptacle fiat, alveolate. Ray-flowers 16-20, pistillate, fertile; ligules
cuneate, 3-cleft. Disk-flowers numerous, hermaphrodite and fertile; corolla-tube densely
glandular, longer than the funnelform throat; teeth lanceolate, longer than the throat.
Achenes elongate and narrowly obpyramidal, 4-angled, striate, rounded at the apex. Pappus
wanting.
Type species, Amauria dissecta A. Gray.
1. Amauriopsis dissecta (A. Gray) Rydberg.
Amauria dissecta A. Gray, Mem. Am. Acad. II. 4: 104. 1849.
Villanova chrysanthemoides A. Gray, PI. Wright. 2: 96. 1853.
Bahia chrysanthemoides A. Gray, Proc. Am. Acad. 19: 28. 1883.
Bahia dissecta Britten, Trans. N. Y. Acad. Sci. 8: 68. 1888.
Eriophyllum chrysanthemoides Kuntze, Rev. Gen. 307. 1891.
Villanova dissecta Rydb. Bull. Torrey Club 37: 333. 1910.
A tall annual; stems 3-6 dm. high, puberulent and glandular, especially above; leaves
1-3 times ternately divided into obovate, oblong, or oblanceolate toothed divisions, puberulent;
involucre hemispheric, 6 mm. high, 10-13 mm. broad; bracts oblanceolate, acuminate.
glandular-hirsute; ligules spatulate, 6-8 mm. long, 3-cleft with rounded teeth; disk-corollas 3
mm. long; achenes narrowly obpyramidal, 4 mm. long, 0.5 mm. thick, glandular-puberulent,
Type locality: A few miles east of Mora River [New Mexico].
Distribution: Wyoming to New Mexico, Arizona, and Chihuahua.
Illustration: Clements, Rocky Mt. Fl. pl. 41, f. 1.
- Baldwin, Bruce G., and Kenneth R. Wood, 2016.
Hymenopappus filifolius Hook. “Fine Leaved Wooly White”
- Hooker, Sir William Jackson, 1829-1840.
• Taxa Notes:
Notes on Hymenopappus filifolius Hook. var. polycephalus;
Original Text
2. H. filifolius ; lana alba decidua tomentosus, ramis paniculatis sparsis, foliis bipinnatifidis
segmentis filiformibus elongatis integerrimis, floribus solitariis, involucri foliolis sub-12
ellipticis pubescentibus capitulo subbrevioribus, corollae tubo snperne campanulato, pappo
brevissimo. — H. tenuifolius. Dougl. MSS. apud. Herb. Hort. Soc. Lond. (non Pursh.)
Hab. On the undulating arid grounds of the Columbia, near the Wallawallah, and on the hanks of the
Spokan and Flat-head Rivers.
—
18 inches to two feet high, the stem loosely panicled above. This is quite
different from the H. tenuifolius of Pursh, of Mhich I possess a specimen from Mr. Nuttall, gathered on the
Arkansa. That has a remarkably dense corymb of flowers, whereas here the branches form a lax terminal
panicle, each elongated extremity bearing a solitary flower. The pappus is extremely minute.
• Golden Checklist Flora:
Hymenopappus filifolius Hook. var. polycephalus;
• Taxa Notes:
Notes on Hymenopappus filifolius, Hooker, 1833;
Hymenopappus filifolius Hook. var. polycephalus (Osterh.) B.L. Turner "Many-Headed Fine-Leaved Wooly-White"
- Osterhout, George E., 1918.
Original Text
Comments
A NEW HYMENOPAPPUS FROM COLORADO
By Geo. E. Osterhout
Hymenopappus polycephalus sp. nov.
Perennial from a tap-root; stems one to several, 4-6 dm. high,
tomentose, becoming glabrate, leafy at the base and less so to the
middle, the stem having 6-8 leaves; much alternately branched,
the branches beginning near the base; leaves 10-12 cm. long,
pinnately or bipinnately parted into very narrowly linear and
rather distant divisions, the upper smaller; inflorescence paniculate,
the heads long peduncled and single at the ends of the
branches; the bracts narrowly obovate, 5-6 mm. long, tomentose,
somewhat thickened in the middle, with a rather broad scarious
margin; flowers yellow, the throat of the corolla broadly campanulate,
scarcely 1.5 mm. long, the reflexed lobes about a third
as much, the tube of the corolla about as long as the throat,
glandular, the pappus about half the length of the tube; the
achenes narrowly obpyramidal 4 mm. long, long villous, but the
pappus not covered by their villosity.
Another perennial, yellow-flowered, Hymenopappus has been
described from Colorado, H. cinereus Rydb., but that is a smaller
plant, with fewer heads of flowers, and is less leafy. Dr. Rydberg's
description says: "Stem about 2 dm. high, branched, with
2-4 leaves." The close relationship of Hymenopappus polycephalus,
however, is not with H. cinereus but with H. tenuifolius Pursh.
It is as high and as leafy as H. tenuifolius, but the
flowers are yellow, not dull white, and it is a perennial, not a
biennial. The achenes are very similar to those of H. tenuifolius.
H. cinereus is a synonym of H. filifolius var. cinereus I.M.Johnst.
H. tenuifolius is an accepted name, for a mostly plains species, with just one collection in Jefferson County.
H. polycephalus is found on the foothills of the eastern side of
the mountains of northern Colorado at an altitude of 6,000 to
7,500 feet. It is plentiful in the open country about Livermore,
Larimer Co., Colo., and northward to Dale Creek, and on into
Wyoming, I think. It blossoms from the last of June to early
in September. I have collected it a number of times, and for a
time thought it belonged with H. tenuifolius. The type specimens
were collected in the vicinity of Livermore, Larimer Co.,
Colo., Aug. 10-11, 1917, No. 5680.
Windsor Colo.
- Turner, Billie L., 1956.
Original Text
Osterhout proposed the specific name polycephalus in 1918,
but it was not taken up by other workers. He apparently was
very familiar with the plant in the field, and his numerous collections
preserved at the Rocky Mountain Herbarium, with
their annotated comments, indicate that he was dissatisfied
with placing it either under H. cinereus Rydb. or H. tenuifolius.
Nevertheless, Osterhout thought polycephalus was more closely
aligned with H. tenuifolius since, in describing the species, he
stated, “The close relationship of Hymenopappus polycephalus,
however, is not with H. cinereus but with H. tenuifolius Pursh.
It is as high and as leafy as H. tenuifolius, but the flowers are
yellow, not dull white, and it is a perennial, not a biennial.”
Johnson treated the entity as a questionable synonym of H.
cinereus stating, “Hymenoppapus polycephalus is a puzzling
form which suggests a perennial phase of H. tenuifolius.”
In the present treatment, polycephalus has been treated as a
variety of H. filifolius since its total characteristics indicate
a much closer relationship to the members of this species. In
spite of this closer affinity, var. polycephalus does have several
characters which possibly link it to H. tenuifolius; indeed,
there seems to be a considerable degree of intergradation between
the two entities with respect to leaf dissection, amount
of tomentum on the stem, and throat/lobe ratio.
It is interesting to note that var. polycephalus, so far as known,
is tetraploid with n = 34, while both var. cinereus and H.
tenuifolius are diploid with n = 17. Var. polycephalus, in
Colorado, occupies a geographical position between these two
taxa. From these facts it might be speculated that at some
past time the Rocky Mountain var. cinereus came in contact
with H. tenuifolius so that extensive hybridization occurred
on the diploid level and concomitantly amphiploidy occurred.
(Evidence of a once farther eastward extension of H. filifolius
var. cinereus is found in the disjunct races of this variety in
the panhandle region of Texas, a considerable distance from
the normal range now occupied by the variety.) With recession
of this peripheral contact at a later time, the amphiploids survived
in the habitats in which they were produced and have
since spread to the area they now occupy. In the few meiotic
slides prepared of var. polycephalus pairing appeared normal,
there being 34 bivalents. Unfortunately, seed of this variety
was germinated too late in the study of this group to provide
material for genetical experiments to test this hypothesis.
• Notes on Colorado Flora:
Leucanthemum vulgare;
Leucanthemum vulgare (Vaill.) Lam. “Ox-eye Daisy”
• Golden Checklist Flora:
Liatris ligulistylis;
Liatris ligulistylis (A. Nelson) K. Schum. “Northern Plains Gayfeather”
• Golden Checklist Flora:
Liatris punctata;
Liatris punctata Hook. “Dotted Blazing Star”
• Golden Checklist Flora:
Lygodesmia juncea;
• Taxa Notes:
Notes on Stephanomeria pauciflora;
Lygodesmia juncea (Pursh) D. Don ex Hooker “Rush Skeletonplant”
• Notes on Colorado Flora:
Madia glomerata;
Madia glomerata Hook. “Mountain Tarweed”
- Hooker, Sir William Jackson, 1829-1840.
2. M. glomerata;
capitulis (2-4) glomeratis paucifloris in corymbam terminalem dispositis, radiis obsoletis.
Hab.
Plains of the Saskatchawan.
Drummond.
—
Aware as I am that Madia viscosa is liable greatly to
vary, I still cannot bring myself to consider the present as a state of that species.
The inflorescence is a true compound corymb,
the branches, which are leafy, bearing at the extremity a cluster of 3-5 few-flowered
capitula, much narrower than those of M. viscosa, and apparently quite destitute of ray.
This is the first time that any Madia has been found on the east side of the great chain
of mountains which separates eastern from western America.
- Nuttall, Thomas, 1840-1841.
* AMIDA.
(A transposition of Madia, in allusion to its affinity to that genus. )
Capitulum heterogamous or homogamous, few-flowered;
rays none, or one or two, irregular, very small, ligulate, three-toothed.
Discal florets hermaphrodite, cylindric, one to five.
Stigma included, small and obtuse.
Receptacle naked, very small.
Involucrum oblong or ovate, of two to five oblong,
or lanceolate sepals, embracing the deciduous fruit.
Achenium oblong, compressed, four-sided, naked, and eranulated ;
in the radial florets sheathed with the sepals.
—
Viscidly glandular annuals, with entire leaves, the lower ones opposite ;
flowers small, in terminal clusters, bracteolate, bractes and
sepals covered with conspicuous, pilose glands.
Allied to Madia, and remarkable for its singular depauperation.
Amida * gracilis; hirsute and scabrous, with close-pressed hairs;
sepals convex, very glandular.
Hab.
Rocky Mountains and prairies of the Oregon.
The whole plant fragrant from glandular exudation.
About a foot high.
The stem simple, sometimes branching towards the summit, slender and rigid.
Leaves narrow-linear, rather crowded, hirsute, and scabrous,
the hairs close-pressed.
Flowers small, yellow, in irregular axillar and terminal clusters;
involucrum sometimes with only one or two flowers, at other times with five.
Rays often wanting.
Amida * hirsuta;
hirsutely pilose with spreading hairs; leaves linear, scabrous, and ciliated on the margin;
sepals hirsute and glandular, carinated.
Hab.
With the above, from which it is distinguished by its pubescence and larger capituli with
broader sepals. The rays are also larger and more regular in their association.
• Golden Checklist Flora:
Mulgedium pulchellum;
Mulgedium pulchellum (Pursh) G. Don in R. Sweet “Blue Lettuce”
- Moulton, Gary E., 1999.
- Pursh, Frederick, 1814.
Original Text
pulchellus.
9. S. pedunculis squamosis,
floribus corymboso-racemosis,
foliis caulinis cordato-amplexicaulibus ovato-oblongis acutis integerrimus glabris.
On the banks of the Missouri.
♃
Sept.
v. s.
Flowers large, beautiful blue.
- Bremer, Kare, 1994.
- Flora of North America Editorial Committee, eds., 2006.
- Weber, William A., and Ronald C. Wittmann, 2012.
- Ackerfield, Jennifer, 2015.
- POWO, 2021 - 2026.
• Golden Checklist Flora:
Nothocalais cuspidata;
Nothocalais cuspidata (Pursh) Greene “Prairie False Dandelion”
- Pursh, Frederick, 1814.
Original Text
Comments and Interpretation
Troximon cuspidatum.
—
T. scapo unifloro superne subtomentoso,
foliis linearibus margine undato tomentosis,
calycinis foliolis imbricatis cuspidatis glabris.
- Gray, Asa, 1884.
• Taxa Notes:
Notes on Comandra umbellata ssp. pallida, DeCandolle, 1857;
Original Text
Comments and Interpretation
§
4. Nothocálais.
Pappus of 20 to 24 narrowly linear-lanceolate silvery-white
paleae, occupying two or more series, with obscure mid-nerve, very gradually
attenuate into a slender awn : akenes attenuate-fusiform : seed not reaching
to the tapering summit : bracts of the oblong-campanulate involucre narrowly
lanceolate, nearly equal, in about two series : perennial from a thick caudex.
Intermediate between Microseris and Troximon !
—
Gray, Proc. Am. Acad. xix. 65.
M. troximoides, Gray.
Acaulescent or nearly so: leaves tufted on the caudex, rather
fleslhy, narrowly linear-lanceolate, entire or undulate, 4 to 6 inches long : scapes a span to a
foot high: involucre three-fourths inch high: ligules somewhat elongated: mature akenes
half-inch long : pappus somewhat longer, its almost setiform paleae a quarter of a line wide
below.
—
Proc. Am. Acad. ix. 211.
—
Wooded hills and open plains, Montana and Idaho
(first coll. by Spalding), Washington Terr., and Oregon to N. W. California.
Another first collection by Spalding was
Comandra umbellata ssp. pallida, also in Washington Territory.
- Greene, Edward L., 1886.
Original Text
N. CUSPIDATA.
—
Akene little contracted, 3 lines long,
filled by the seed : pappus of 40 — 50 unequal, very narrow,
setose paleae and scabrous bristles: leaves all radical, longer
than the flowering scapes: involucre glabrous.
—
Troximon Pursh, Fl. ii. 742; Torr. & Gray, Fl. ii. 489; Gray, Syn. Fl. ii. 437:
T. marginatum, Nutt. Gen. ii. 127.
Oligosporus Cassini “Tarragon, Sagewort”
- Cassini, Alexandre H. G., 1817.
• Glossary:
calathid;
synantheree;
• Taxa Notes:
Notes on Artemisia campestris, Weber & Wittmann, 2012;
Notes on Artemisia dracunculus, Weber & Wittmann, 2012;
Original Text
Translation
51. Oligosporus.
Ce genre, ou sous-genre, de la tribu des anthemidées,
comprend toutes les espčces d' Artemisia, L., dont la calathide est
composée de fleurs femelles et de fleurs males.
Telle est, par exemple, l' Artemisia campestris, L.
51. Oligosporus.
This genus, or subgenus, of the anthemid tribe,
includes all species of Artemisia, L., whose calathid is
composed of female and male flowers.
Such is, for example, Artemisia campestris, L.
Packera A. Löve & D. Löve
There are no collections of P. streptanthifolia made in Jefferson County.
- Harrington, H. D., 1964, 2nd ed..
- Löve, Áskell, and Doris Löve, 1976.
- Weber, W. A., and Áskell Löve, 1981.
- Trock, Debra, 2006.
Filename: 280Packera_plattensis_streptanthifolia.html
Packera streptanthifolia (Greene) W. A. Weber & Á. Löve, Phytologia. 49: 48. 1981.
Packera pseudaurea (Rydberg) W. A. Weber & Á. Löve, Phytologia. 49: 48. 1981.
Packera plattensis (Nuttall) W. A. Weber & Á. Löve, Phytologia. 49: 48. 1981.
Packera fendleri (A. Gray) W.A. Weber & Á. Löve
Coll. No. 2064
Rocky Mountain groundsel
False-gold groundsel
Prairie groundsel
Fendler's Ragwort
Perennials, 10–50+ cm; fibrous-rooted (caudices weak to stout, horizontal to suberect).
Perennials, 20–70+ cm; fibrous-rooted (bases simple or branched, horizontal to erect).
Biennials or perennials, 20–60+ cm; rhizomatous and/or fibrous-rooted (bases erect to suberect), sometimes stoloniferous (mostly eastern populations).
Perennials, 10–40+ cm; rhizomatous (rhizomes horizontal to suberect, branched).
Perennial herb, 25-35 cm., caudex , some, perhaps older ones, are stout and erect;
Stems 1 or 2–5, clustered, usually glabrous, sometimes sparsely floccose-tomentose proximally and in leaf axils.
Stems usually 1, sometimes 2–4, clustered, glabrous or sparsely tomentose proximally.
Stems 1 or 2–3, clustered, floccose-tomentose proximally and in leaf axils, otherwise sparsely tomentose or glabrescent.
Stems 1 or multiple (crowded to subcespitose), floccose-tomentose or glabrescent.
Stem, tomentose, especially in axils, thinning distally;
Basal leaves (and proximal cauline, relatively thick and turgid) petiolate; blades spatulate to oblanceolate, or ovate to orbiculate, 20–40+ × 10–30+ mm, bases tapering to abruptly contracted or subcordate, margins entire, crenate, dentate, or weakly lobulate (faces usually glabrous, sometimes hairy).
Basal leaves petiolate; blades usually broadly lanceolate to ovate, sometimes subhastate, 20–50+ × 20–40+ mm, bases truncate to subcordate or obtuse, margins crenate, denticulate, bluntly dentate, or sharply dentate (proximal cauline leaves petiolate; margins usually pinnately lobed to laciniate, sometimes subentire).
Basal leaves (and proximal cauline) petiolate; blades narrowly elliptic to elliptic-ovate or oblanceolate to suborbiculate or sublyrate, 20–70+ × 10–30+ mm, bases tapering to rounded or abruptly contracted, margins subentire to crenate, serrate-dentate, or pinnately lobed (abaxial faces floccose-tomentose, especially along midribs, ± glabrescent).
Basal leaves petiolate; blades lanceolate to oblanceolate, 30–60+ × 10–30+ mm, bases tapering, margins shallowly, evenly pinnatifid to pinnatisect or wavy (adaxial faces floccose-tomentose or subglabrescent).
Basal leaves, petiole, 35-40 mm, long, thin not winged, tomentose, blade 20-24 mm. × 7-9 mm. wide, ovate, some (7 of 23 = 30%) lyrate, 2-7-2.8 × longer than wide, mid-vein tomentose thinning distally, tips, mostly entire, some 3-toothed;
Cauline leaves gradually to abruptly reduced (± petiolate or sessile; entire or subentire).
Cauline leaves gradually reduced (becoming sessile, sometimes clasping).
Cauline leaves gradually reduced (petiolate, sublyrate or pinnatisect, abaxial faces sparsely hairy; distals sessile, subentire to irregularly dissected).
Cauline leaves gradually reduced (sessile; lanceolate to oblanceolate, pinnatisect to wavy).
Cauline leaves, reduced distally, sessile, base not auriculate, inreasingly pinnatisect distally;
Heads 2–20+ in loose, corymbiform or subumbelliform arrays. Peduncles bracteate, glabrous or sparsely tomentose.
Heads 5–20+ in open or congested, subumbelliform arrays.
Heads 6–20+ in open or congested, corymbiform arrays. Peduncles conspicuously bracteate, sparsely to densely tomentose.
Heads 6–25+ in open or compact, corymbiform arrays.
Inflorescence, heads #3-5 per stem, >leaves;
Peduncles bracteate, glabrous.
Peduncles bracteate, densely to irregularly floccose.
Peduncles, 8-12 mm.;
Calyculi conspicuous.
Calyculi inconspicuous.
Calyculi inconspicuous.
Calyculi 0 or inconspicuous (bractlets red-tinged).
(Calyculi inconspicuous)
Involucre, 5 mm. × 9 mm. wide, ovoid, thinly tomentose;
Phyllaries (8–)13 or 21, green (tips sometimes cyanic), 4–7+ mm, glabrous.
Phyllaries (13–)21(–30+), light green, 3–8 mm, glabrous.
Phyllaries 13 or 21, green (tips sometimes cyanic), 5–6+ mm, densely tomentose proximally, glabrescent distally.
Phyllaries 13, green, 5–7 mm, floccose proximally to glabrescent distally.
Phyllaries, in 2 equal series, 5.5 mm. × 1.0-1.3 mm. wide, green, thinly tomentose, margins, flat, scarious, tip, reddish brown;
Receptacle, epaleate;
Flowers, of 2 kinds;
Ray florets 8 or 13; corolla laminae 5–10 mm.
Ray florets 0, 8, or 13; corolla laminae 6–10+ mm
Ray florets 8–10; corolla laminae 9–10 mm.
Ray florets 6–8+; corolla laminae 5–7 mm.
Ray flowers, #10-12, tube 3 mm. + blade 6.5-7.5 mm. × 2.7 mm. wide, color yellow, fertile;
Disc florets 35–60+; corolla tubes 2–4 mm, limbs 2.5–4 mm.
Disc florets 70–80+; corolla tubes 2.5–3.5 mm, limbs 2–3 mm.
Disc florets 60–70+; corolla tubes 2.5–3.5 mm, limbs 3.5–4.5 mm.
Disc florets 30–40+; corolla tubes 2.5–3 mm, limbs, 2.5–3.5 mm.
Disk flowers many, tube 4 mm. + lobes 0.5 mm., throat expanding gradually, yellow, open, bisexual;
Pappus, many, well-developed. bristles, 4.5 mm., ±equal;
Cypselae 1–2.5 mm, glabrous; pappi 3–6 mm.
Cypselae 1–1.5 mm, glabrous; pappi 4.5–5.5 mm.
Cypselae 1.5–2.5 mm, usually hirtellous, sometimes glabrous; pappi 6.5–7.5 mm.
Cypselae 2.5–3 mm, glabrous; pappi 4–5 mm. 2n = 46.
Fruit, 1.2 mm. × 0.5 mm. wide, compressed front-to-back, color brown, margin, thinly pubescent;
2n = 46, 92.
2n = 46, 92.
2n = 46.
Flowering late May–late Aug. Forests, open meadows, valleys, dry to damp and loamy soils; 1000–3400 m; Alta., B.C., N.W.T., Sask., Yukon; Calif., Colo., Idaho, Mont., Nev., N.Mex., Oreg., Utah, Wash., Wyo.
Flowering mid Apr–early Jun(–mid Jul, north). Prairies, meadows, open wooded areas, along highways, railroads, around mining and construction areas, usually on limestone; 50–1800 m; Ont., Sask.; Ark., Colo., Ga., Ill., Ind., Iowa, Kans., La., Mich., Minn., Miss., Mo., Mont., Nebr., N.Mex., N.Dak., Ohio, Okla., Pa., S.Dak., Tenn., Va., Wis., Wyo.
Flowering late May–early Oct. Steep slopes, loose, dry rocky or gravelly soils, along streams, open forests, disturbed sites; 1600–3200 m; Colo., N.Mex., Wyo.
Packera streptanthifolia is widespread and variable throughout the Western Cordillera. It includes weakly defined phases that have been treated as distinct species or as varieties. Characteristics used to delimit those taxa often overlap and are difficult to score; some ""phases"" grade into each other. Northern populations are sometimes segregated as a distinct taxon (e.g., Senecio streptanthifolia var. borealis; J. F. Bain 1988).
Packera plattensis is abundant, widespread, and almost weedy. Putative hybrids with other species are known. Plants in mesic, remnant prairies in the east are sometimes stoloniferous.
Packera fendleri is abundant, almost weedy in the southern Rocky Mountains. It thrives in a wide range of elevations and in a wide variety of habitats; flowering times vary. It frequently grows in close association with other species of Packera and may hybridize with them.
- Weber, William A., and Ronald C. Wittmann, 2012.
Original Text
Interpretation
3a. Basal leaves and most stem leaves deeply pinnatifid or runcinate-pinnatifid.
P. fendleri (A. Gray) Weber & Love.
Abundant in gravelly soil, open forests of the foothills.
Producing rosettes from long, slender rhizomes; leaf lobes uniform, shallow; leaves white tomentose.
3b. Basal leaves oval, rately pinnatifid except at the very base. … 4
4b. Basal leaves narrower, on winged petioles, irregularly toothed, lobed or entire. … 10
Leads eventually to P. plattensis.
Original Text
Interpretation
2a. Basal and stem leaves nearly all deeply and evenly pinnately dissected or runcinate-pinnatifid … 3
This couplet leads to P. multilobata and P. fendleri.
2b. Basal leaves with entire, toothed, or crenate margins
(in P. plattensis and P. tridenticulata a few basal leaves may have pinnatisect margins,
but the majority of the leaves will have entire or merely toothed margins),
stem leaves entire to pinnatisect or sublyrate … 4
This couplet eventually leads to P. plattensis after elininating seven other taxa,
and arriving at a group including P. plattensis, P. streptanthifolia, and P. pseudaurea.
• Golden Checklist Flora:
Packera fendleri;
Packera fendleri (A. Gray) W.A. Weber & Á. Löve. (Syn: Senecio fendleri A. Gray ) Fendler"s Ragwort.
- Gray, Asa, 1849.
• Publication Details:
Gray, 1849, publication details;
Original Text
Comments
444. S. Fendleri (sp. nov.) :
perennis, flocoso-incanus, demum subglabratus ;
caule folioso erecto pedali corymbosi-ramoso ;
foliis oblongis omnibus pinnatifidis supra glabratis inferioribus in petiolum nudum attenuatis summis sessilibus haud amplexicaulibus, segmentis 11-21 confertis oblongis obtusissimis plerisque inciso-dentatis seu 2-4-lobatis ;
corymbis compositis polycephalis ;
involucro campanulato fere ecalyculato 12-phyllo multifloro ;
ligulis 7-8 oblongis disco duplo longioribus ;
acheniis glaberrimis.
—
Foot of mountains along the Creek, twelve miles above Santa Fe ;
June, July.
(478, † 480.)
—
A well-marked species, related to S. eremophilus.
Stem stout.
Leaves from 2 to 4 inches long including the petiole,
some of the lower often only sinuate-pinnatifid,
but commonly all deeply pinnatifid or pinnately parted,
the crowded lobes from one fourth to half an inch long, early glabrate above ;
but still floccose of white-wooly underneath.
Involucre 3 lines long.
Rays 4 or 5 lines long. *
• Golden Checklist Flora:
Packera plattensis;
Packera plattensis (Nuttall) W. A. Weber & Á. Löve. “Prairie Groundsel”
- Nuttall, Thomas, 1840-1841.
• Publication Details:
Nuttall, 1841, publication details;
Original Text
Senecio * Plattensis;
♃,
somewhat pubescent;
base of the stem arachnoidly tomentose;
leaves all pinnatifid, the radical petiolate, cauline amplexicaule,
lobes oblong, denticulate, the centre lobe sublanceolate;
corymb nearly simple;
involucrum subcampanulate, minutely bracteolate;
sepals about twenty, scute;
rays usually twelve, oblong, a little longer that the short involucrum;
achenium puberulous;
pappus about the length of the florets.
Hab.
In the Rocky Mountain range, and in Arkansa.
About ten to fourteen inches high ;
stem simple, striated.
Corymb nearly simple, with ten to twelve heads of flowers,
pedicels one to two inches long, slightly bracteolate.
The Arkansa specimen is taller and more slender,
with the primary small radical leaves entire and smooth,
the leaves more elongated, and less denticulate.
The whole habit of the plant, as well as the flowers,
are very similar to S. tomentosus,
at lease the smoother variety, but the achenium is less pubescent.
• Golden Checklist Flora:
Packera tridenticulata;
Packera tridenticulata (Rydb.) W.A. Weber & Á. Löve “Three-Tooth Ragwort”
- Gray, Asa, and Charles Wright, 1852.
- Rydberg, Per Axel, 1900b.
• US Highway 24:
in Buena Vista;
Original Text
Comments
7. Senecio tridenticulatus sp. nov.
Senecio aureus var. borealis A. Gray, Pl. Wright. I : 125. 1852 ;
Not T. & G. 1843 ;
S. aureus var. compactus A. Gray, Syn. Fl. I2: 391 ; in part.
Perennial with a branched caudex, in age perfectly glabrous,
or slightly floccose at the base of the leaves : stems very slender,
about 3 dm. high : basal leaves very narrowly oblanceolate, thick
with a slender petiole, slightly 3-toothed at the apex or wholly
entire, 4-8 cm. long and 4-5 mm. wide: stem leaves linear and
subentire : cyme open and corymbiform : heads 7-8 mm. high
bracts lanceolate, acute ⅔ or ¾ as long as the disk : rays light
yellow, about 8 mm. long and 1—1.5 mm - wide > 3-nerved ; achenes
hispid-puberulent on the angles.
This species differs, from the preceding in the slender stem and
the open cyme, and also in the form of the leaves. The latter
character also separates it from the next following. It grows in
wet sandy soil. The type was growing at an altitude of 2400 m.
Wright's specimens are past blooming, and good characters could
not be taken from them, wherefore I have made Sheldon's speci-
mens the type. The latter were mixed with some of the next.
[Plate 5, f. 12.]
The preceding was Senecio compactus (A. Gray)Rydb.
and the following was Senecio oblanceolatus Rydb.,
both of which are now treated as synonyms of Packera thurberi (A.Gray) B.L.Turner.
Colorado : Cottonwood Creek, Buena Vista, 1892, C. S. Sheldon
(type in the herbarium of N. Y. Botanical Garden).
Texas: Mountains beyond the Limpia, 1849. Wright, 403.
See Plantae Wrightianae (Gray & Wright, 1852).
- Weber, W. A., and Áskell Löve, 1981.
• Notes on Colorado Flora:
Packera werneriifolia;
Packera werneriifolia (A. Gray) W. A. Weber & Á. Löve “Hoary Groundsel”
• Notes on Colorado Flora:
Pericome caudata;
Pericome caudata A. Gray “Mountain Tail-Leaf”

Pericome caudata A. Gray “Mountain Tail-Leaf” along the Peaks to Plains Trail.
• Publication Details:
Gray, 1853, Plantć Wrightianć, Part II;
Pericome caudata.
—
Among rocks, on the sides of mountains, at the copper
mines, New Mexico ; Oct. (1195.)
—
“
Plant 4 feet high, much branched, growing
in large bunches.”
Base of the stem not gathered. Branches terete, striate, sparingly puberulent.
Leaves nearly all opposite, on long petioles, membranaceous,
3-nerved at the base, dilated triangular, or the upper hastate, with the basal angles
usually acuminate or more or less produced, often angulate-toothed below the middle,
the apex produced into a very long tail-like acumination.
The larger leaves in
the specimens are 3 inches broad at the nearly truncate base, and 4 inches long,
including the acumination, which is 1½ to 2 inches in length ; the uppermost are
narrower, but equally caudate. They are sparingly puberulent, especially on the veins,
sparsely sprinkled or dotted with minute resinous atoms, like a Eupatorium ; but
not at all pellucid-glandular after the mode of Tagetinese. Heads in terminal
cymes or corymbs, on slender pedicels, half an inch long. Involucre not bracteate,
about 3 lines long ; the scales very narrowly linear, one-nerved, their narrowly scarious
margins lightly but decidedly coalescent to near the apex, the free tips
lanuginous-ciliate.
Corolla with the elongated and exserted anthers 3 lines long, yellow ;
the slender tube very viscous-glandular, about the length of the cylindraceous
throat and short limb. Branches of the style convex externally, slightly flattened,
the rather indistinct stigmatic lines extending to very near the summit, and there
insensibly vanishing, where a fine pubescence begins ; the same pubescence extends
down the back considerably lower, and disappears insensibly. Achenia 2 lines
long, much compressed, margined all round with a strong callous nerve, which
bears a dense and conspicuous beard of hispid hairs. Pappus rather shorter than
the beard of the achenium, composed of numerous setiform squamellae which are
irregularly united below into a hyaline lacerate crown. There is no trace of any
awns.
—
A genus of even more doubtful position in the family than Perityle, Benth.,
which with Laphamia, I have continued to append to the Asteroideae. It agrees
with Perityle in the achenia and the (disk) corollas, in the prevailingly opposite
leaves, and very nearly in the style ; but difl'ers in the absence of a ray, as well as
of awns to the pappus, and in the coalescence of the strictly uniserial scales of the
involucre. The style would almost do for the Eupatoriaceae ; but the stigmatic
lines are continued almost to the tip ; and the flowers are yellow. The involucre
is like that of Hymenatherum, and the style is exactly that of a Tagetes; but
there are no large pellucid glands, and the achenia are flat. Notwithstanding, I
should append the genus to Tagetineae, were it not for the many points of
resemblance to Perityle.
• Golden Checklist Flora:
Picradeniopsis oppositifolia;
Picradeniopsis oppositifolia (Nutt.) Rydb. ex Britton. “Opposite Leaf Bahia”
- Moulton, Gary E., 1999.
- Nuttall, Thomas, 1818.
Original Text
Comments
568. * TRICHOPHYLLUM. †
Calix oblong-cylindric, many-leaved, equal.
Radial florets oblong.
Receptacle naked.
Pappus paleaceous, minute, 5 to 8-leaved, leaflets obtuse, awnless.
Herbaceous;
leaves alternate? or opposite, palmately pinnatifid, tomentose or villous;
peduncles 1-flowered, dichotomal and terminal.
Species.
1. T. lanatum.
Actinella lanata, Ph. 2. p. 560.
Every where whitely and lanuginously tomentose;
leaves alternate,
those of the stem subpalmately pinnatifid, of the branches linear and entire;
peduncle elongated, the summit thicker.
Hab.
Near the sources of Columbia river.
M. Lewis.
Flowering in June and July.
v. s. in Herb. Lambert.
—
Perennial; stem erect and branching, about a foot high.
Leaves alternate? (perhaps not constantly so)
those of the stem elongated, narrow at the base,
dilated and divided pinnatifidly above, division ligulate and somewhat toothed, uppermost entire.
Calix oblong-cylindric, composed of a simple series of leaves,
about 12 to 14, linear-lanceolate, acute.
Rays about the same number, oblong, bidentate.
Pappus 5 to 8-leaved.
Seed pentangular? glabrous, attenuated downwards.
—
The flowers are bright yellow, and in form and character strongly
resemble those of the genus Tagetes.
This would become Eriophyllum lanatum (Moulton, 1999).
2. * oppositifolium.
Decumbent and much branched, shortly and canescently pubescent;
leaves opposite, all palmately trifid, segments ligulate, simple or divaricately subdivided;
peduncle filiform, mostly dichotomal, scarcely longer than the leaves.
Hab.
On denudated sterile hills, near Fort Mandan; abundant.
Flowering in July and August.
—
Perennial? stem diffuse, 6 to 12 inches high, grooved; oppositely branched.
Leaves petiolate, trifid, canescent, pubescence very short,
segments about an inch long, thickish and opaque,
the lateral ones mostly bifid, the central one trifid,
all somewhat obtuse and linear.
Peduncle slender, 1 to 2 inches long, a little thicker under the calix.
Calix oblong-cylindric, simple, leaflets 5 to 8,
oblong-ovate, erect;
rays about the same number, very short.
Pappus paleaceous, 5 to 8-leaved, minute, leaflets partly obtuse and somewhat lacerate.
Seen nearly smooth, rather long, and attenuated downwards, or inversely conic.
Receptacle small and naked.
—
The whole of this plant is very sensibly bitter and destitute of aroma.
There is nothing in the habit of this genus which would lead us to
suppose it allied to Actinella of Jussieu,
and scarcely more in the generic character.
In Actinella the calix is very short, flat, and horizontally spreading;
the leaflets of the paleaceous pappus awned, and the seeds villous;
the leaves are also alternate and entire.
The proximate affinity of the present genus is to Tagetes.
† The copious pubescence,
particularly distinguishing these plants from the genus Tagetes.
- Britton, Nathaniel Lord, 1901.
Original Text
Comments
76. PICRADENIOPSIS Rydb.
[Bahia Nutt., not Lag.]
Britton gives Rydberg credit for the genus name, but I don't see a place that Rydberg ever published it himself.
… [English genus diagnosis not reproduced.] …
[Name referring to resemblance of this genus to Picradenia.]
1. Picradeniopsis oppositifolia (Nutt.) Rydb. False Bahia. (I. F. f. 3967.)
…. [English description not reproduced.] …
On plains, S. Dak., Neb. and Mont. to Kans. and N. Mex.
June-Sept.
[Bahia oppositifolia Nutt.]
Reference is to the “Illustrated Flora” figure 3967.
• Golden Checklist Flora:
Prenanthes racemosa;
Prenanthes racemosa Michx. “Purple Rattlesnake Root”
• Golden Checklist Flora:
Pseudognaphalium canescens;
Pseudognaphalium canescens (DC.) Anderberg “Wright's Rabbit-Tobacco”
- DeCandolle, Augustus Pyramus, 1836.
Original Text
Translation or Comments
40. G. canescens, totum villis sericeo-lanatis canum, canle herbaceo erecto ramoso,
foliis lineari-lanoeolatis acuminatis , infer. basi attenuatis, super. breviter
adnatis , capitulis ad apices ramulorum congestis sessilibus , involucre
ovato-campanulati squamis scariosis albis lanceolatis acuminatis disco aequalibus.
♃
in Mexico ad Leonem ultra Guanaxuato legit cl. Mendez.
Folia 1 ½ poll. longa, 2-3 lin. lata, patula.
Inv. circ. 25-phyllum. Flor. herm. 5-7, foem. circ. 25.
(v. s.)
León, Guanajuato, Mexico collected by Mendez.
Seen in the dried state by DeCandolle.
- Anderberg, Arne A., 1991.
• Golden Checklist Flora:
Pseudognaphalium macounii;
Pseudognaphalium macounii (Greene) Kartesz “Macoun's Rabbit-Tobacco”
- Greene, Edward L., 1902b.
Gnaphalium macounii. Apparently biennial, the stems
rigidly erect, about two feet high, rather loosely leafy and clothed
with a somewhat hirsute and viscid glandular-pubescence : leaves
narrowly oblanceolate, acute, 3 inches long, the upper decurrent,
all white-woolly beneath, light green and merely glandular-pubescent above :
branches of the subpyramidal close panicle and the
main stem for some distance below it densely white-woolly :
involucres of middle size, their pearly scarious bracts all ovate, very
acute : flower and fruit not seen.
Collected in the Chilliwack Valley, B.C., 29 July, by Mr. Jas.
M. Macoun, No. 26,847 ; also earlier at Revelstoke, No. 11,334,
and again from the Warm Springs, Kootenay Lake, both in
British Columbia, in the year 1890. No. 34,053 from Salmon
Arm, J. R. Anderson, 1899, is also the same. The species is related
to G. decurrens, yet very distinct in habit and inflorescence, the
dense white-woolly pubescence of the upper part of stems and
branches of the panicle being very peculiar.
Ratibida Raf. “Prairie Coneflower”
- Rafinesque, C. S., 1818b.
125. Rudbeckia columnaris must form the genus Ratibida of Raf. fl. Miss.
• Golden Checklist Flora:
Ratibida columnifera;
• Water Tank Road:
above the curve;
• Field Notes:
Coll. No. 1198, 12 Jul 2015;
Ratibida columnifera (Nutt.) Woot. & Standl. “Upright Prairie Coneflower”
[So, is the cypsela ciliate on the abaxial side, or the adaxial side?]

Coll. No. 1198, Ratibida columnifera
- Fraser, John, 1813.
- Greene, Edward L., 1889.
• Glossary:
flosculi;
fulvous;
• Publication Details:
Fraser's Catalogue, publication details;
Original Text
Comments
Tagetes patula is commonly called the French Marigold, also in the Asteraceae, and native to Mexico and Guatemala.
I think this might be what we now call Echinacea purpurea, not native to Colorado,
but widely cultivated, and found occasionally as a garden escapee.
* New Species. — † A Shrub. — ‡ Perennial. — M. from Missourie.
It had passed through the hands of Nuttall, and had received one or two slight corrections from his pen.
Moreover, he had written his name in ink, as the author of the Catalogue …
- Pursh, Frederick, 1816.
• Publication Details:
Pursh, 1814, publication details;
Original Text
657. RUDBECKIA. Gen. pl. 1324.
…
11.
R. caule stricto simplici summutate paucifloro,
pedunculis elongatis, foliis pinnatifidis incisis,
laciniis linearibus,
calyce simplici 5-phyllo, radiis 5-8.,
disco cylindraceo elongato.
R. columnifera, Fraser catal. 1813
On the Missouri. v. s.
The singular appearance of the receptacle of this plant
distinguishes it from all the other known species.
- Fraser, John, 1813.
- Nuttall, Thomas, 1818.
• Publication Details:
Nuttall, 1818, publication details;
Original Text
590. RUDBECKIA. L.
Calix subequal, mostly consisting of a double series of leaflets.
Receptacle paleaceous, conic.
Pappus a 4-toothed margin.
Herbaceous; leaves alternate, entire, lobed of pinnaifid ;
flowers terminal ; disk often dark, rays yellow,
rarely brown, in R. purpurea purple.
— Stigma often obtuse.
Species.
....
12. columnaris.
Hispid; stem nearly simple, 1 or few-flowered,
peduncles very long;
radical leaves nearly entire, cauline pinnatifid,
segments linear-lanceolate;
calix simple, 5 to 8-leaved, rays 5 to 8;
disk cylindric, elongated.
Hab.
On the plains of Upper Louisiana.
Flowering in July.
Perennial: 1 to 2 feet high.
Rays sometimes brown-red, as in Tagetes patula.
13.
...
A North American genus,
with the exception of R. nudicaule of Monte Video,
which appears to be scarcely distinct from R. spathulata.
The seeds of R. purpurea are pungently aromatic.
- Rafinesque, C. S., 1818a.
- Rafinesque, C. S., 1818b.
• Publication Details:
Amer Monthly Mag Crit Rev, publication details;
Ratibida columnaris Raf. is treated as a synonym of R. columnifera (Nutt.) Wooton & Standl.
Original Text
125. Rudbeckia columnaris must form the genus Ratibida of Raf. Fl. Miss.
- Wooton, E. O., and Paul C. Standley, 1915.
Original Text
2.Ratibida columnifera (Nutt.) Woot. & Standl.
Rudbeckia columnifera Nutt. Fraser's Cat. No. 75. 1813.
Rudbeckia columnaris Pursh, Fl. Amer. Sept. 575. 1814.
Ratibida columnaris D. Don in Sweet, Brit. Flower Gard. II. 4: pl. 361. 1838
Lepachys columnaris Torr. & Gray, Fl. N. Amer. 2:313. 1842.
2a. Ratibida columnifera pulcherrima (DC.) Woot. & Standl.
Obeliscaria pulcherrima DC. Prodr.5:559. 1836.
Ratibida columnaris pulcherrima D. Don in Sweet, Brit. Flower Gard. II. 4: pl. 361. 1830
Lepachys columnaris pulcherrima Torr. & Gray, Fl. N. Amer. 2: 313. 1842.
This is a mere form of the type and hardly deserves a name.
Both forms almost invariably occur together,
although occasionally they grow alone,
It is possible to find in a single patch every possible gradation
in the color of the rays from pure bright yellow to solid brown-purple.
The same variation in color occurs in R. tagetes,
but since that has very small and inconspicuous rays no one has yet
thought to distinguish the various forms by name.
- Reveal, James L., 1968.
• Publication Details:
Reveal (1968) comments on Fraser (1813) Catalogue - 1;
75.
Rudbeckia columnifera Nutt. In Fras. Catal. 1813.
In my opinion, this species is adequately described in Fraser's Catalogue, as Nuttall states:
“Spontaneous varieties of this plant sometimes occur
with bright fulvous flowers, coloured like Tagetes patula:
the stem is simple, seldom producing more than three flowers,
which are of an uncommon length,
appearing like a column of flosculi,
subtended by 5-8 neutral florets,
and a simple calyx.”
This species was also described as R. columnaris Sims, Bot. Mag. 39: 1601. 1813,
from cultivated plants given to Sims by the Fraser Brothers
who had grown the species from seeds collected by Nuttall.
The Pursh name, R. columnaris Pursh, Fl. Am. Sept. 2: 575. 1814,
is an illegitimate substitute for Nuttall's R. columnifera and is thus an exact synonym of it.
The species is now commonly known as
Ratibida columnifera (Nutt. In Fras.) Woot. & Standl., Contr. U.S. Nat. Herb. 19: 706. 1915.
- Richards, Edward Leon, 1968.
• Notes on Colorado Flora:
Rudbeckia hirta;
Rudbeckia hirta L. “Black-Eyed Susan”
• Notes on Colorado Flora:
Rudbeckia hirta pulcherrima;
Rudbeckia hirta L. var. pulcherrima Farw. “Black-Eyed Susan”
• Notes on Colorado Flora:
Rudbeckia laciniata;
Rudbeckia laciniata L. “Cut-Leaf Codeflower”
- Linne´, Carl von, 1753.
RUDBECKIA
laciniata.
1. RUDBECKIA foliis compofitis lcainiatis.
Vir. cliff. 88.
Hort. cliff. 420.
Hort, upf. 269.
Roy. lugdb. 181.
Gron. virg. 180.
Chryfanthemum americanum perenne, foliis divifis dilatius virentibus, majus.
Morif. hift. 3. p. 22. f. 6. t. 6. f. 53. & 54.
Doronicum americanum, laciniato folio.
Baub. pin. 516.
Aconitum helianthemum canadenfe.
Corn. canad. 178. t. 179.
Habitat in Virginia, Canada.
♃
• Notes on Colorado Flora:
Rudbeckia laciniata ampla;
Rudbeckia laciniata L. var. ampla (A. Nelson) Cronquist “Cut-Leaf Coneflower”
- Nelson, Aven, 1901.
Rudbeckia ampla
Perennial, from horizontal rootstocks with large fleshy crowns
from which spring numerous semi-fleshy roots: stems clustered,
i. e., several from the same tuft, 1-2 m. high, glabrous, strongly
striate, moderately branched above: leaves numerous and ample,
light green and glabrous below, a shade or two darker above with
minute scattering hispid hairs from pustulate bases (the hairs sometimes
deciduous), with conspicuous, light-colored veins: basal
leaves large, 2-3 dm. broad, trifoliate, on petioles 2-4 dm. long:
the leaflets petiolulate, broadly oval or ovate in outline, three-cleft,
the segments coarsely and irregularly serrate, the middle segment
often again lobed or cleft : lower stem-leaves similar but smaller,
the petioles becoming slenderer and shorter upward, the leaves
merely 3-parted: upper stem-leaves sessile, 3-parted or merely
cleft, 1-2 dm. long ; the segments becoming entire as to margin,
the lateral oblong-lanceolate, the middle variously toothed and
cleft: rameal leaves reduced, becoming entire and bract-like, from
broadly oval-acute to lanceolate: heads terminating the branches,
large : involucral bracts oblong to ovate, obtuse or subacute, 8-12
mm. long: rays 6-12, conspicuous, 3—5 cm. long, 8-15 mm. wide:
disk in full anthesis and in fruit cylindrical-ovate, 2-4 cm. high.
That this is in part the R. laciniata of Gray’s Syn. Fl. 262 is
possible but it is far from the typical eastern R. laciniata L. The
pinnately arranged narrow segments with wide sinuses of the
smaller leaves of that, need but to be compared with the trifoliate-palmate
broad segments and narrow sinuses of this to see that
they are distinct. The larger rougher disk, the fewer and stouter
peduncles and the different involucral bracts are equally characteristic of R. ampla.
Whether all the Rocky Mountain forms belong here I am
unable to say, but I would include
Baker’s no. 699, Pagosa Springs, Colo.;
224, Jennie M. Archibald, Berwind, Colo.;
and all my numbers as follows:
Eagle Rock Canyon, Aug., 1891; no. 1575,
Cotton Wood Canyon, Aug. 5, 1895;
no. 7709, Centennial, July 26, 1900.
- Hitchcock, C. Leo, et al., 1955.
Senecio accedens Greene
- Greene, Edward Lee, 1896.
• Taxa Notes:
Notes on Senecio bigelovii hallii, Greene 1896;
Senecio accedens.
Stems several from the root, about
2 feet high, leafy below, somewhat corymbosely panicled
above, with rather few and large nodding heads: leaves
narrowly lanceolate, acute, denticulate, on long and slender
petioles which are much dilated below, the whole herbage
sparsely flocculent when young, at flowering time glabrous
or nearly so: heads about inch high; bracts of the narrow-campanulate
involucre oblong-lanceolate, scarious-margined,
the few calyculate ones ligulate; rays none ; ovaries glabrous.
• Taxa Notes:
Notes on Senecio bigelovii hallii;
Senecio bigelovii A. Gray “Bigelow's Ragwort”
- Torrey, John, and Asa Gray, 1857a.
Senecio Bigelovii (sp. nov.) :
glabra ;
caule simplici e radice perenni apice racemoso-3-15-cephalo ;
foliis ovato-lanceolatis acuminatis argute calloso-dentatis in petiolum longum marginatum contractis,
supremis lanceolatis basi augusta semiamplexicaulibus ;
capitulis magnis nutantibus homogamis ;
involucro late campanulato 10-12-phyllo basi bracteolis paucis brevibus setaceis calyculato,
squamis acutis aequalibus, exterioribus lanceolatis, interioribus latioribus
scarioso-marginatis ;
ligulis nullis ;
acheniis glaberrimis.
In mountain arroyos, near Camp Douglas, New Mexico ; October.
Plant entirely glabrous.
Stem rather stout, erect, 18 inches to 2 feet or more in height, rather leafy to the top ;
the uppermost leaves reduced to bracts.
Lower leaves 3 to 5 inches long, abruptly contracted at the base into a margined or winged
petiole of 2 or 3 inches in length ;
the upper successively narrower and with shorter petioles, or
at length sessile.
Heads racemose, nodding on the summit of erect and naked or slightly
bracteolate peduncles of 1½ to 3 inches in length, very large for a Senecio,
from half to three quarters of an inch in length and breadth, many-flowered.
Involucre rather fleshy, a little shorter than the flowers, very minutely bracteolate ;
the scales 5 or 6 lines long, herbaceous,
with abrupt hyaline-scarious margins, which on the alternate and interior scales are broad and
conspicuous.
Corollas yellow, narrowly infundibuilform-tubular,
rather deeply 5-toothed at the summit, externally callous- thickened, and reticulated at the tip.
Stamens, style, &c, as in Senecio.
Achenia linear, perfectly glabrous, strongly ribbed or angled, fully 3 lines long.
Pappus soft and fine, white, minutely scabrous.
This striking species may perhaps be taken for a Cacalia ;
but I see nothing to distinguish it from Senecio.
There is no North American species with which I can compare it, except S. Fremontii, Torr. & Gray,
which has much smaller and radiate heads.
• Notes on Colorado Flora:
Senecio bigelovii hallii;
• Taxa Notes:
9330510280Senbig;
Senecio bigelovii A. Gray var. hallii A. Gray “Hall's Ragwort”

Coll. No. 3629, Senecio bigelovii var. hallii.
Georeferenced vouchers of Senecio bigelovii var. hallii in CCH2 and SEINet.
- Gray, Asa, 1863.
• Taxa Notes:
Notes on Senecio bigelovii hallii, Nuttall, 1841;
…
321. S. Bigelovii, var. Hallii. ║ “Subalpine; heads very drooping, rayless.”
…
║ Senecio Bigelowii,
(Gray in Bot. Whippl. Exped. Pacif. R. R. Surv. 4, p. (55) 111), var. Hallii:
foliis fere omnibus lanceolatis cum caule pilis articulatis pubescentibus(demum glabratis), caulinis
omnibus sessilibus imisve in petiolum alatum contractis. S. megacephalus, Nutt., thus far found
only by Nuttall, has a similar pubescence, but more of it,and also on the involucre; the scales
of the latter are narrower, the heads are radiate and erect, and the plant is dwarf.
- Greene, Edward Lee, 1896.
• Taxa Notes:
Notes on Senecio accedens, Greene, 1896;
- Rydberg, Per Axel, 1917.
1. S. accidens Greene. Stem 4-10 dm. high, simple and leafy; basal leaves
and lower stem-leaves tapering into rather long winged petioles, lanceolate or
oblanceolate, 10-15 cm. long, dentate or denticulate; upper stem-leaves
linear-lanceolate, sessile and somewhat clasping; heads few or several on erect branches,
nodding; involucre 2-2.5 cm. broad; bracts glabrous, thin, oblong-linear, acute.
S. Bigelovii Hallii A. Gray. S. scopulinum Greene. Mountains: Colo. — Utah
— N.M. — Wyo. Mont. Jl-O.
• Golden Checklist Flora:
Senecio eremophilus var. kingii;
Senecio eremophilus Richardson var. kingii (Rydb.) Greenm. “Cut-Leaved Groundsel”
- Rydberg, Per Axel, 1910b.
Original Text
Senecio Kingii sp. nov.
Senecio eremophilus D. C, Eat., Bot. King's Exp. 191. 1871.
Not S. eremophilus Richards. 1823.
Perennial, with a thick rootstock; stems glabrous, 3-6 dm.
high, rather stout, leafy; leaves obovate or oblanceolate In outline,
4-7 cm. long, the lower petioled, all pinnately lobed one
third to one half the distance to the midrib, with ovate or lanceolate,
more or less toothed lobes; heads numerous, corymbose-paniculate,
9-11 mm. high; involucres glabrous, campanulate,
7-8 mm, high, 6-7 mm. broad; bracts linear, acute, carinate,
sometimes with small black tips; the calyculate ones few, subulate;
rays 5-7 mm. long; achenes scabrous-papillose on the rounded
angles.
This species is related to S. eremophilus, but differs in the
somewhat smaller and erect heads, less deeply dissected leaves, and
their broad and short divisions, and shorter rays. One of the
specimens cited below was determined some years ago as S. glaucifolius,
but that species differs from this as well as from the rest
of the group in the narrower and scarcely carinate bracts.
Utah: Cottonwood Canon, Aug. 1869, S. Watson 676 (type,
in herb. Columbia University); Alta, Wahsatch Mountains, 1879,
M. E. Jones 1144; American Fork Canon, July 1895, M. E. Jones;
Big Cottonwood Canon, Aug. 1905, Garrett 1591; near Marysvale,
1905, Rydberg & Carlton 7068; Mount Barrette, 7206; Fish Lake,
7206.
• Golden Checklist Flora:
Senecio integerrimus;
Senecio integerrimus Nutt. “Columbia Ragwort”
• Glossary:
carnose;
sphacelate;
• Publication Details:
Nuttall, 1818, publication details;
Original Text
Comments
564. SENECIO. L. (Ragwort.)
Calix cylindric, subcaliculate:
scales sphacelate at the point.
Receptacle naked.
Pappus simple, capillary, and copious.
sphacelate = undergo necrosis.
Suffruticose or more commonly herbaceous;
leaves entire or pinnatifid;
flowers mostly corymbose or terminal;
yellow or rarely purple.
A few species are destitute of rays.
Species. 1. …
15. * integerrimis
Smooth; stem simple and attenuated;
leaves perfectly entire;
radical ones long petiolate, lanceolate, acute, cauline, sessile,
acuminate, uppermost minute;
corymb simple, 8 to 12-flowered? peduncles 1-flowered,
rays shorter than the hemisperical caliculate calix.
HAB.
In depressed and moist situations on the plains of the Missouri, near the Great Bend.
Flowering in June.
Flower large and yellow.
Stem 12 to 18 inches high.
Lower leaves thickish and somewhat carnose,
very smooth, uppermost minutes, slightly tomentose;
corymb coarctate.
Seeds smooth.
Nearly allied to S. aquaticus.
carnose = of a fleshy consistence —used of succulent parts of plants.
A genus of more than 140 species principally indigenous to Europe and the Cape of Good Hope.
- Cronquist, Arthur, 1950.
Senecio integerrimus Nutt. var. exaltatus (Nutt.) Cronquist, comb. nov.
S. exaltatus Nutt. Trans. Am. Phil. Soc. II, 7:410 (1841)
S. lugens Richards. var. exaltatus A. Gray, Bot. Calif. 1:413 (1876).
Senecio megacephalus Nuttall “Nuttall's Ragwort”
- Nuttall, Thomas, 1840-1841.
• Taxa Notes:
Notes on Senecio bigelovii hallii, Gray, 1863;
Senecio * megacephalus;
♃,
stem low, densely lanuginous at base; leaves deciduously lanuginous,
oblong-lanceolate, entire, cauline amplexicaule, linear-lanceolate, acute;
corymb of three to five bracteolate, large capituli; involucrum pubescent,
of twenty to twenty-four linear, acute sepals, tipped with tufts
of hairs; rays pale yellow, ten to twelve, scarcely as long as the involucrum;
achenium smooth, with about ten striatures; dentures of the florets papillose.
Hab..
On the plains of the Platte, towards the Rocky Mountains. Nearly allied to S. alpestris.
About six to eight inches high; the leaves perfectly entire, more or less pubescent beneath, lower
leaves with long petioles, Capituli very large, for the size of the plant, about as large as those
of the common bur, (Arctiwm lappa.)
• Golden Checklist Flora:
Senecio riddellii;
Senecio riddellii (Torr. & A. Gray) Greenm. ex L.O. Williams “Riddell's Ragwort”
• Golden Checklist Flora:
Senecio spartioides;
• Field Notes:
6 Sep 2020;
Senecio spartioides Torr. & A. Gray. “Broomlike Ragwort”

Coll. No. 2231, Senecio spartioides with a praying mantis and a beetle.
Senecio spartioides adventive in my garden.
- Torrey, John, and Asa Gray, 1838-1843.
• US Highway 287:
at Jeffrey City;
Original Text
* * * * Perennial : heads corymbose, chiefly radiate
† Leaves entire or denticulate.
8. S. spartioides :
glabrous throughout;
stems suffruticose, very numerous from the same ligneous tap-root, rigid,
corymbose at the summit, leafy;
leaves fleshy, narrowly linear, perfectly entire,
rather obtuse, sessile;
heads (large and showy) fastigiate-corymbose,
on short minutely bracteolate peduncles ;
the calyculate scales subulate, minute ;
scales of the cylindrical involucre about 12, lanceolate-linear, acutish;
rays mostly 7, oblong-linear, elongated ;
achenia silky-canescent.
Upper Platte;
on a steep sand-bank of the Sweet-water River,
Lieut. Fremont!
Aug.–Sept. — Stems a foot high, forming a dense tuft.
Leaves 1-3 inches long, about a line wide, very numerous.
Heads half an inch in length.
Rays golden-yellow.
Pappus as long as the disk-corolla.
—
A remarkable and handsome species.
• Golden Checklist Flora:
Senecio wootonii;
Senecio wootonii Greene “Wooton's Ragwort”
Solidago L. “Goldenrod”
- Semple, J. C., 2018.
• Golden Checklist Flora:
Solidago canadensis;
Solidago canadensis L. “Canada Goldenrod”
• Golden Checklist Flora:
Solidago gigantea;
Solidago gigantea Aiton “Giant Goldenrod”
Original Text
Comments
4. S. caule erecto glabro, foliis lanceolatis glabris ferratis margine feabris,
racemis paniculatis fecundis,
pedunculis hirtis, ligulis abbreviatis.
gigantea.
4. Solidago smooth stiff stems,
leaves are smooth, margins serrate;
panicles with clusters of flowers;
peduncles rough, ligules short.
Gigantic Golden-rod.
Nat. of North America.
Cult. 1758, by Mr. Philip Miller.
Mr. Philip Miller was the curator at the Physick Garden at Chelsea.
H.
♃
H. – Hardy.
♃. – Perennial.
• Golden Checklist Flora:
Solidago missouriensis;
Solidago missouriensis Nutt. “Missouri Goldenrod”

Habitat of Coll. No. 1256, Solidago missouriensis at Ranson/Edwards.
Coll. No. 1732, Solidago missouriensis at North Table Mountain.
- Nuttall, Thomas, 1834a.
• Publication Details:
Nuttall, 1834a, publication details;
Even though this was described with collections by Wyeth,
the locations noted seem much more like Nuttall collections
from his 1811 trip on the Missouri River and 1819 trip to Arkansas.
Wyeth collections would have been along the Platte, Snake, or Columbia.
Original Text
55. Solidago * Missouriensis.
Pumila, glabra, recemis erectis, foliis lineari-lanceolatis, acutis,
inciso-subserrularis, superioribus integris,
panicula brevi laxa, floribus majus-culis
Stem slender, smooth, leafy, about a foot or so high.
Leaves scabrous at the margin.
Panicle about three inches long, the branchlets slender,
the flowers pedicellate,
and brought together in a somewhat rhomboidal raceme.
Rays as long as the calyx.
Hab.
in the upper branches of the Missouri and in Arkansas.
• Golden Checklist Flora:
Solidago nana;
Solidago nana Nutt. “Baby Goldenrod”
- Nuttall, Thomas, 1840-1841.
• Taxa Notes:
Crepis occidentalis, Weber & Wittman, 2012;
• Publication Details:
Nuttall, 1841, publication details;
Original Text
Solidago * nana;
somewhat cinereous and pulveulently pubescent, dwarf,
many stems from the same root;
lower and radical leaves spathulate, obtuse, entire,
or subserrulate at the apex, stem leaves linear, narrowed below;
ramuli fastigiate, subcorymbose;
bractes linear;
involucrum nearly smooth, scales ovate;
rays about seven, oblong, as long as the disk;
achenium pubescent.
HAB.
In the Rocky Mountain range, near Lewis' River of the Shoshonee.
About a span high,
with a large, black, almost woody root.
Stem leaves small,
radical ones about one and a half inches by half an inch wide;
scales of the involucrum unusually broad,
pubescent on the margin,
rays conspicuous.
Apparently allied to S. nemoralis, though very distinct and alpine.

Coll. No. 2439, Solidago nana.
• Golden Checklist Flora:
Solidago nemoralis ssp. decemflora;
Solidago nemoralis Aiton ssp. decemflora (de Candolle) Brammall ex Semple “Gray Goldenrod”
Original Text
Translation
15. S. decemflora, caule erecto tereti foliisque pube brevissima subscabris,
foliis oblongis mucronulatis integerrimis subtriplirierviis ,
radicalibus basi attenuatis,
racemis subsecundis in paniculam corymbosam dispositis,
invol. glabri squamis linearibus , capitulis 10-floris, ligulis 5 minutissimis.
♃ in Mexicanae prov. Texas districtibus orientalibus legit cl. Berlandier
(pl. exs. n. 1924).
Achaenia sericeo-villosa. Herba pedalis. (v. s.)
15. Solidago decemflora stems erect,
leaves rounded and short scabris;
oblong leaves mucronulate entire subtri-nerved;
Radical base attenuate,
corymbose panicle, subsecundis in the clusters have been arranged,
involved, smooth, linear phyllaries, heads 10-flowered, ligules 5 minute.
♃ (perennial) In Mexican Prov. East Texas, eastern districts
per the collector Berlandier
(Pl. Exs. N. 1924).
Achaenia silky-villous. The plant a foot tall.
(Seen in the dried condition.)
• Golden Checklist Flora:
Solidago simplex;
Solidago simplex Kunth “Sticky Goldenrod”
- Kunth, Carol Sigismund, 1815-1825.
Original Text
Translation and Comment
1. SOLIDAGO SIMPLEX. †
S. caule erecto, simplici, subangulato, striato, superne hirtello; foliis lineari-lanceolatis,
superne obsolete crenatis, inferne valde angustatis, glabris , margine denticulato-scabris ;
panicula terminali, ramis subtrifloris.
Crescit prope pagum Sanctce Rosae Mexicanorum , alt. 1300 hex. ♃ Floret Septembri.
It grows near the Mexican village called Santa Rosa, altitude 1300 hex.
perennial. Flowers in September.
Radix crassa, perpendicularis, fibris crebris obsita.
Caulis erectus, i 4 — i 5 -pollicaris , simplex,
striatus, subangulatus, puiqrarascens, superne hirtellus.
Folia radicalia sessilia , lineari-laneeolata ,
acutiuscula , inferne valde angustata , adeo ut petiolata appareant, subintegerrima aut apicem versus
remote et obsolete crenata, reticulato-venosa, crassiuscula, glabra, margine denticulis minutissimis
scabra, 26—27 lineas longa , tres lineas lata. Folia caulina alterna , radicalibus simillima, magis
tamen acuta , summa subacuminata. Panicula terminalis, subramosa , bracteata5 ramis subternis, patulis,
subtrifloris, angulatis , hirtellis; floribus pedicellatis , magnitudine floris Solidaginis Virgaureae;
bracteis linearibus.
Involucrum campanulato-hemisphaerieum; foliolis circiter 25 , laxe imbricatis , linearibus,
acutis , planis , uninerviis , margine scariosis et diaphanis , glabris, nitentibus; exterioribus
brevioribus.
Receptaculum convexum, scrobiculatum, glabrum.
Flosculi disci circiter 20, tubulosi
hermaphroditi; radii 8— g, ligulati, feminei. Flosculi hermaphroditi :
Corolla flava, glabra, superne
ampliata , quinquefida ; laciniis lineari-lanceolatis , acutis, enerviis , patulis.
Stamina tubo corollae inserta.
Antherae subinclusae.
Ovarium lineare , hispidum.
Stylus glaber.
Stigma bipartitum, exsertum;
laciniis patentibus , glabriusculis.
Akenium immaturum.
Pappus pilosus, sessilis; pilis scabris, apice vix
incrassatis, albis, corollam subsequantibus, patulis.
Flosculi feminei:
Corolla flava ; tubo tenui, glabro;
ligula lanceolato-oblonga, tridentata, plana, patente , sesquilineam longa.
Ovarium, Stylus, Stigma et Pappus prorsus ut in flosculis hermaphroditis.
Pappus tubo paulo longior.
Solidagini Virgaureae habitu affinis.
Solidago virgaurea has similar habitat.
• Golden Checklist Flora:
Solidago speciosa;
Solidago speciosa Nutt. “Showy Goldenrod”
- Nuttall, Thomas, 1818.
• Publication Details:
Nuttall, 1818, publication details;
Original Text
560. SOLIDAGO. L. (Golden-Rod.)
...
†† Racemes erect
27. *speciosa.
Stem tall and smooth, simple or virgately branched;
leaves lanceolate, entire, somewhat carnose.
scabrous on the margin, the lower very broad,
radical ones subserrate;
racemes mostly terminal, erect and compound, pubescent;
pecuncles mostly shorter than the calix;
rays elongated about 5;
seed smooth.
Hab.
In shady woods, on the banks of the Schuykill, also in Jersey;
near Philadelphia, but rare.
S. sempervirens. Mich.
S. integrifolia? Persoon, 2. p. 449.
Allied to S. petiolaris.
Stem often 6 feet high, smooth and sulcate.
Lowest leaves a span long, and 3 inches broad,
irregularly and remotely subserrate, upper leaves very entire.
gradually diminishing upwards, in dry and shady situations,
membranacei=ous and veined,
in gardens subcarnose and smaller, with the veins partly obliterated,
racemes also numerous, but always rigid, terminal and erect.
Flowers larger than the preceding, with the calix also coloured
(which in the preceding is green);
rays bright yellow, unusually broad.
The seeds in this species are perfectly smooth,
in our sempervirens pubescent.
This is one of the most ornamental plants of the genus.
• Golden Checklist Flora:
Solidago velutina ssp. sparsiflora;
Solidago velutina DC. ssp. sparsiflora (A. Gray) Semple. “Three-Nerve Goldenrod”
- Gray, Asa, 1876a.
- Wheeler, George M., 1878.
Solidago sparsiflora. Virgaurea, Virgatae : scabrido-puberula ;
foliis inferioribus ignotis, su; erioribus floralibusque parvulis lanceolatis
(lin. 6-12 longis); racemulis oligocephalis laxis laxeque thyrsoideis ;
involucri squamis linearibus puberulis apice viridulis acutiusculis ;
floribus radii circiter 10 ligulis parvulis, disci 4-5 ;
acheniis sericeo-pubescentibus.
— Arizona, near Camp Lowell, Sept., 1874. Rothrock, in
Wheeler Expedition, 1874.
• Golden Checklist Flora:
Sonchus arvensis;
Sonchus arvensis L. “Field Sowthistle”
• Golden Checklist Flora:
Sonchus oleraceus;
Sonchus oleraceus L. “Common Sow-Thistle”
• Golden Checklist Flora:
Stephanomeria pauciflora;
• Taxa Notes:
Notes on Lygodesmia juncea;
Stephanomeria pauciflora (Torr.) A. Nelson “Brownplume Wirelettuce”
- Torrey, John G., 1828.
Original Text
Comments
202. Prenanthes juncea, Pursh fl. ii. p. 498.
Nutt. gen. ii. p. 123. On the Platte and Missouri.
This is a synonym of Lygodesmia juncea (Pursh) D. Don ex Hooker
Obs. Near the Rocky Mountains, Dr. James collected two
species of a genus allied to Prenanthes, but differing in their
plumose and sessile pappus. They will probably hereafter
be referred to a new genus. Their principal characters are
as follows:
1. P.? pauciflora, caule ramoso, flexuoso, superne paniculato,
ramulis unifloris, floribus erectis ; foliis lineari-lanceolatis,
runcinatis, glabris ; calycibus quinquefloris.
This will become Stephanomeria pauciflora (Torr.) A.Nelson in J.M.Coult. & A.Nelson,
New Man. Bot. Centr. Rocky Mt. 588 (1909)
2. P.? tenuifolia, caule ramoso, glabro ; foliis linearibus,
integerrimis; panicula laxa terminali; floribus erectis;
calycibus quinquefloris.
This is a synonym of Stephanomeria tenuifolia (Torr.) H.M.Hall,
of which most Colorado collections are on the western slope.
- Coulter, John M., and Aven Nelson, 1909.
Original Text
Comments
2. Stephanomeria pauciflora (Torr.) Nels. Similar to the preceding, usually
lower, smaller in all of its parts, and somewhat woody at the base: the lower
leaves at least more or less runcinate-pinnatifid: pappus plumose nearly to
the base. S. runcinata in part. (Ptiloria pauciflora Raf. Atl. Journ. 145.
1832.) Sparingly if at all in our range; on the plains; from Nebraska to Texas.
The preceding was S. tenuifolia (Torr.) Hall.
• Golden Checklist Flora:
Symphyotrichum ericoides;
• Publication Details:
Dill. elt., 1732;
Gron. virg., 1738;
Roy. lugdb., 1740;
Symphyotrichum ericoides (Linnaeus) G. L. Nesom “White Heath Aster”
- Linne´, Carl von, 1753.
Original Text
Translation and Comments
16. ASTER caule paniculato, pedunculis racemofis : pedicellis foliofis :
foliolis linearibus integerrimus.
Gron. virg. 100.
ericoides
16. ASTER stem paniculate, peduncles racemose : pedicels leafy :
leaves linear entire.
After caule paniculato, floribus racemofis, calycibus patulis oblongis,
foliis fubulato-linearibus. Roy. lugdb. 168.
Aster with paniculate stems, flowers recemose, calyx wide oblong,
leaves subulate-linear.
After ericoides dumofus. Dill. elt. 40. t. 36. f. 40.
Aster ericoides dumosus.
Habitat in America feptentrionalis.
♃
Lives in North America.
- Nesom, Guy L., 1994.
• Golden Checklist Flora:
Symphyotrichum falcatum;
Symphyotrichum falcatum (Lindl.) G.L. Nesom “White Prairie Aster”
• Golden Checklist Flora:
Symphyotrichum fendleri;
Symphyotrichum fendleri (A.Gray) G.L.Nesom “Fendler's Aster”
- Gray, Asa, 1849.
• US Interstate 25:
at Canadian R;
at Ocate Ck;
†
317. A. Fendleri (sp. nov.):
caulibus spithamaeis e caudice sublignosa pluribus
adscendentibus rigidis parce hispidulis ;
ramis corymboso-paniculatis monocephalis ;
foliis sessilibus rigidis coriaceis parvis linearibus mucronulatis integerrimis glabris
uninerviis marginibus hispido-ciliatis, infimis subspathulatis, ramealibus brevissimis ;
involucri campanulati squamis 3-seriatis lineari-oblongis glanduloso-scabrellis mucronulatis,
exterioribus herbaceis obtusis laxis, interioribus acutis paulo longioribus;
acheniis pubescentibus. —
†
317. A. Fendleri (sp. nov.):
stems with many spithamae from a sublignous caudex
ascending rigid, sparingly hispidulis;
branches with corymbose-paniculate, monocephalic;
leaves with sessile, rigid, leathery, small, linear, mucronulate, entire, glabrous
with single-nerved, hispido-ciliate margins, the lowest subspathulate, very short rameals;
involucres with campanulate scales with 3-seriate linear-oblong, glandular-scabrellious,
mucronulate,
exterior herbaceous, obtuse, loose, interior acute, slightly longer;
achenes pubescent. —
On the Ocaté Creek and the Rio Colorado (Upper Canadian) ;
Aug. (372.) —
On the Ocaté Creek and the Rio Colorado (Upper Canadian) ;
Aug. (372.) —
A low species, of the group Ericoidei, or perhaps Amelli,
with much the aspect of Diplopappus linariifolius.
The longer leaves are only three fourths of an inch in length.
Involucre one fourth of an inch in diameter.
The rays have apparently been violet or blue.
A low species, of the group Ericoidei, or perhaps Amelli,
with much the aspect of Diplopappus linariifolius.
The longer leaves are only three fourths of an inch in length.
Involucre one fourth of an inch in diameter.
The rays have apparently been violet or blue.
• Notes on Colorado Flora:
Symphyotrichum foliaceum parryi;
Symphyotrichum foliaceum (DC.) G. L. Nesom var. parryi (D. C. Eaton) G. L. Nesom “Parry's Aster”
• Golden Checklist Flora:
Symphyotrichum laeve;
Symphyotrichum laeve (L.) Á. Löve & D. Löve var. geyeri (A. Gray) G.L. Nesom “Smooth Blue Aster”
• Golden Checklist Flora:
Symphyotrichum lanceolatum ssp. hesperium;
Symphyotrichum lanceolatum (Willd.) G. L. Nesom ssp. hesperium (A. Gray) G. L. Nesom Western Lance-Leaved Aster
- Gray, Asa, 1884.
Original Text
A. hesperius.
Resembles A. paniculatus and A. salicifolius of the East, equally variable,
from nearly glabrous and smooth to scabrous-pubescent . leaves lanceolate, entire or the
larger with a few denticulations (2 to 5 inches long, 3 to 8 lines wide) : heads rather
crowded, 4 or 5 lines high : involucre of narrowly linear or more attenuate acute or gradually
acuminate erect bracts, either unequal and imbricated, or with some loose and slender
herbaceous exterior ones which equal the inner : rays either white or violet, 3 or 4 lines
long. — Damp soil and along streams, S. Colorado and New Mexico to Arizona
and S. California.
Has been variously taken for A. longifolius, Novi-Belgii,
aestivus, &c., and coll. by
Wright, Greene, Rothrock, Cleveland, Parish, Lemmon, &c.
• Golden Checklist Flora:
Symphyotrichum porteri;
Symphyotrichum porteri (A. Gray) G.L. Nesom “Smooth White Aster”
- Porter, Thomas C., and John M. Coulter, 1874.
Original Text
Aster ericoides, L., var. strictus, Porter. Low, ¾°-1° high, glabrous,
except the scabrous margins and ciliate bases of the leaves, erect,
slender, paniculately branched above, branches short; scales of the involucre
narrowly linear, lax, outer ones very acute, often entirely green,
inner ones scarious with a central green line; radical leaves narrowly
oblanceolate. — “In the mountains at middle elevations,”
Hall & Harbour, 254. Near Denver, Coulter.
Foot-hills west of Denver, Porter; Meehan; Hoopes.
- Gray, Asa, 1880.
Among the true Asters are several forms which have to be named,
such as A. Porteri for A. ericoides, var. strictus, Porter & Coult. Fl.
Colorad. 56, …
- Semple, J. C., and J. G. Chmielewski, 2025.
• Golden Checklist Flora:
Taraxacum officinale;
Taraxacum officinale F. H. Wigg. “Common Dandelion”
• Notes on Colorado Flora:
Tetraneuris ivesiana;
Tetraneuris ivesiana Greene “Ives' Four-Nerve Daisy”
• Golden Checklist Flora:
Thelesperma megapotamicum;
Thelesperma megapotamicum (Spreng.) Kuntze “Hopi Tea Greenthread”
- Sprengel, Curt Polycarp Joachim, 1826.
Original Text
Translation and Comments
454 Syngenesia. II. Eupatorinae. Bidens
454 Syngenesia. II. Eupatorinae. Bidens
megapotamica*
33. B. foliis omnibus 2pinnatifidis lineari-filiformis glabris,
floribus fubgeminis pedunculatis erectis difcoideis,
involucro colorado anthodium aequante.
Ad fl. magnum Amer. auftr.
(Rio grande. Sello.)
megapotamica*
33. Bidens leaves all 2-pinnatifid, linear-filiform, glabrous,
peduncle erect,
heads equal, sub-budded (refers to calyculi??),
involucre reddish-brown,
flowers discoid.
On large rivers of the American west.
(Rio grande. Sello.)
- Kuntze, Otto, 1891-1898.
Original Text
Thelespermum megapotamicum OK.
(Bidens m. Spr. 1826 —
Th scabiosodes Less. 1831.)
Argentina: Villa Merceded, Ceres.
• Golden Checklist Flora:
Townsendia exscapa;
Townsendia exscapa (Richards.) Porter “Stemless Townsend Daisy”
- Hooker, Sir William Jackson, 1829-1840.
Original Text
58. TOWNSENDIA. Nov. Gen.
Capitulum heteroganium.
Flosculi radii ligulati foeminei; disci hermaphroditii regulares.
Involucri foliola plurisermlia imbricata.
Pappus uniserialis; radii brevis, paleaceo-subulatus,
inaequalis; disci pilosus, pilis elongatis scabriusculis.
—
Herba Americae borealis humilis acaulis.
Radix fusiformis, lignosa, longe descendens ; superne saepe divisa, reliquiis foliorum
vetustorum obtecta. Folia omnino radicalia, numerosa, erecto-patentia, linearia, subspathulata,
integerrima, utrinque adpresso-sericea, basi dilatata, et utrinque ciliata.
Flos solitarius, foliis
immersus et obtectus, ratione plantae magnus.
Involucrum ovatum, foliolis sericeis, lineari-subulatis,
erectis, imbricatis.
Flosculi radii ligulati, erecti, pallide rosei, marginibus involuti,
apice dentibus tribus conniventibus.
Stigma bifidum, purpureum, glabrum.
Ovarium oblongum, compresso-triquetrum, pubescens.
Pappus brevis sed valde inaequalis, uniierialis, e pilis seu
paleis subvlatis, basi membranaceis.
Flosculi disci tubulosi, lutei.
Pappus pilosus, pilis
scabriusculis longitudine corollae.
Ovarium sericeum.
Receptaculum alveolato-punctatum, nudum.
1. T. sericea. (Tab. CXIX), Aster? exscapus. Rich, in Frankl.
1st. Journ. ed. 2. App. p. 32.
Hab. Carlton House upon the Saskatchawan. Dr. Richardson.
Rare, upon the dry banks of the Saskatchawan and among the Rocky Mountains. Drummond.
—
This highly interesting little plant, no less on
account of its habit than its pappus, deserves to be separated from Aster, of which,
indeed, it was by Dr. Richardson, considered a doubtful species.
“the bud is formed in the autumn,” and what Dr. Richardson
further observed in the living plant,
I find to be characteristic of all the specimens in this Collection,
that “the florets of the ray are mostly involute, rarely expanded, and always narrow,
nearly of the same colour with the pappus and inconspicuous; the flowers, indeed, n
ever fully expanding.”
I have named the Genus in compliment to David Townsend, Esq.
of West Chester, Pennsylvania, who having imbibed the most ardent love of
Botany from his friend and instructor Dr. Darlington of the same city,
has devoted his leisure hours to the science with eminent success.
The plant now under consideration is peculiarly worthy of bearing his name,
because he has studied and ably discriminated the numerous Pennsylvanian species of the
allied Genus Aster.
Tab. CXIX. — Fig. 1, Flowers ;
fig. 2, Floret of the ray;
fig. 3, Portion of the pappus from the ray;
fig. 4, Floret of the disk;
fig. 5, Portion of a hair from the pappus of the disk;
fig. 6, Stigma from the disk;
fig. 7, 7, Scales of the involucre;
fig. 8:
— all more or less magnified.

Hooker (1834), Tab. CXIX, Townsendia exscapa
Townsendia grandiflora Nutt. “Largeflower Townsend Daisy”
• Golden Checklist Flora:
Townsendia grandiflora;
Original Text
Comments
Townsendia * grandiflora;
stem canescent, divaricately branching from the base,
branches one or few-flowered,
leaves linear-sublanceolate, very acute, nearly smooth,
or minutely pubescent, green ;
capitulum hemispherical ;
involucrum of three series, the sepals lanceolate, filiformly acuminate,
minutely fringed ;
rays twenty-eight to thirty, or more, bidentate.
Hab.
With the preceding, which it resembles wholly in habit,
but with the flower as large nearly as that of the China Aster,
(Callistephus Chinensis)
Branching from the base, and spreading out sometimes from six to ten inches along the ground,
Leaves linear, much attenuated below, and very acute,
when green rather succulent, and appearing smooth,
though somewhat pubescent beneath, (seen through a glass.)
Sepals elegantly imbricated, perfectly lanceolate, much acuminated,
scariose, except the centre, which is green,
the margin minutely lacerate-ciliate.
Rays pale lilac, longer than the disk.
—
A plant which well deserves cultivation, from its large, showy flowers.
The prededing was Townsendia strigosa which was found
“On the Black Hills, (or eastern chain of the Rocky Mountains,)
near the banks of the Platte.” In Colorado, it is found only in westernmost counties.
• Golden Checklist Flora:
Townsendia hookeri;
Townsendia hookeri Beaman “Hooker's Townsend Daisy”
- Nuttall, Thomas, 1840-1841.
Original Text
TOWNSENDIA. (Hooker.)
Townsendia sericea; caespitose; leaves narrow linear, acute, scarcely half a
line wide, canescently sericeous; capituli sessile on the caudex: scales of the
involucrum numerous, very narrow and acuminate. — Achenium as in the rest
of the genus, obovate, margined, and flatly compressed, sericeous, with a
numerous connate series of white, silky pappus, almost plumosely barbellate, and
remarkably attenuated above.
Hab. On the Black Hills, (an alpine chain toward the sources of the Platte.)
Flowering probably in April.
By the achenium, this genus makes some approach to Calimeris, though totally
unlike in habit.
Nuttall goes on the describe Townsendia incana that he describes as with the above.
- Beaman, John H., 1957.
10. Townsendia Hookeri Beaman, sp. nov.
Type: Clokey 4338, dry hills, Mt. Vernon Canon, 1730 m. alt.,
Jefferson Co., Colorado, 13 April 1920 (*COLO 12061, holotype; CAN,
CAS, DS, F, GH, MICH, MO, MONTU, NA, PH, POM, *RM, UC, US, UTC, WS,
WTU, isotypes).
• Notes on Colorado Flora:
Townsendia incana;
Townsendia incana Nuttall “Hoary Easter Daisy”
• Golden Checklist Flora:
Xanthisma spinulosum;
Xanthisma spinulosum (Pursh) D. R. Morgan & R. L. Hartm. “Spiny Goldenweed”

Coll. No. 2438, Xanthisma spinulosum
- Nuttall, Thomas, 1813.
81 * [Sideranthus] pinnatifidus.
Fl. of both these species yellow,
and considerably like those of the genus Aster,
but from which they are sufficiently distinct.
M

Coll. No. 2434, Xanthisma spinulosum
- Pursh, Frederick, 1814.
Original Text
650. AMELLUS. Gen. pl. 131 5.
...
...
spinulosus.
2. A. canescens ; foliis bipinnatifidis inciso-dentatis, laciniis
linearibus rigido-mucronatis, floribus lateralibus et
terminalibus congestis, radiis bidentatis, paleis setaceis.

Detail of Coll. No. 2434, Xanthisma spinulosum
- Moulton, Gary E., 1999.
- Morgan, David R., and Ronald L. Hartman, 2003.
14. Xanthisma spinulosum (Pursh) D. R. Morgan & R. L. Hartman, comb. nov.
Basionym: Amellus spinulosus Pursh, Fl. Amer. Sept. 2:564. 1813.
…
Type: U.S.A. [South Dakota. Lyman or Brule Co.: Reveal et al. 1999], in open
prairies on the Missouri, 15 Sep 1804, M. Lewis s.n. (Lectotype: PH! [designated by
Hall 1928]; isolectotype: UC!).
…
The following two binomials are not valid as the genus Sideranthus was a nomen
nudum when the combinations were published: Sideranthus pinnatifidus Nutt.,
Fraser's Catalogue, no. 81, 1813, non Aplopappus pinnatifidus Nutt., 1840 and
Sideranthus spinulosus (Pursh) Sweet, Hort. Brit., 227. 1826.
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