Tom Schweich  

Source Data

Salix

Types and Namesakes from Mono Lake or Mono County

Eriogonum kearneyi Tidestr. var. monoense (S.Stokes) Reveal

Eriogonum nodosum Small subsp. monoense Stokes, subsp. nov. Perenne, amplum multis ramis intricatis et divaricatis suberectis, praeter fibres omnino dense floccoso-tomentosum ; pedunculis multo et obtuse ramosis ; bracteis parvis, acutis ; foliis late ovatis, 1-3 cm. longis et paulo angustioribus, petiolis brevioribus ; involucris sessilibus, soils in axillis. apice ramulorum subcapitatis et dense confertis, dentibus invertis, majoribus 2 mm. diametro ; floribus 1.5-2 mm. longis, fere inclusis in involucris, basi obtusis, segmentis perianthii obtusis, pallidis.

Type : Herb. Calif. Acad. Sci. No. 298959, collected Aug. 7, 1938, at the summit of the Sherwin Grade, Mono County, California, by J. T. Howell, No. 14362. It was collected also by Mark Kerr, No. 453, on Mt. Whitney Road. 6000 ft. elev. The subspecies is founded upon the small involucres and flowers and the subcapitate arrangement of the involucres (Stokes, 1943).

Polygonaceae Eriogonum kearneyi Tidestr. Var. monoense (S.Stokes) Reveal -- Leafl. W. Bot. 10: 334. 1966 (GCI)

The concept of E. nodosum has, at least mainly in California, been applied to ssp. monoense S. Stokes (described in Leaft. West. Bot. 3: 201, —1943). This taxon seems better treated as E. kearneyi Tidest. var. monoense (S. Stokes) Reveal, stat. & comb. nov. The var. kearneyi ranges from western Utah and adjacent northwestern Arizona westward across central Nevada to Washoe Co., and enters California in the Mono Lake Basin. The var. monoense extends southward from the Sierra Nevada above Owens Valley to the San Bernardino Mountains, and differs in its longer, more clustered involucres and in the larger, more robust habit of the plants (Reveal, 1966).

Polygonaceae Eriogonum kearneyi subsp. monoense (S.Stokes) Munz ex Reveal -- Madroño 25: 61. 1978 (GCI)

Eriogonum nummulare M. E. Jones in the Jepson Manual, 1st Edition.

Thalictrum monoense Greene

Ranunculaceae Thalictrum monoense Greene -- Leafl. Bot. Observ. Crit. 2. 93 (1910). (IK) F. V. Coville & F. Funston, 1806, 1891-07-08, Locality: Cottonwood Creek, White Mountains, a synonym of Thalictrum alpinum L..

THALICTRUM MONOENSE. Plants slender, 3 to 5 inches high ; foliage short and compact ; leaflets only 11, mostly as broad as long, obtusely about 3-lobed, dull glaucescent green above and there mostly veinless except as to the lobes, these marked by 1 to 3 slender but sharply prominent whitish veins, the lower face very glaucous and venulose : flowering pedicels short, slender, pendulous : sepals oblong-lanceolate, acutish, thin, slightly purple-tinted : stamens about 6 ; ovaries 3 or 4, stipitate : fruit not seen.

White Mountains, Mono County, California, 12 July, 1891, Coville & Funston, No. 1806 ; type specimen on National Herbarium sheet No. 294. Doubtless related to T. duriusculam of Utah, but upper face of foliage differing as to hue and venation, also dull-green, whereas that of the Utah species is distinctly lustrous as in T. alpinum (Greene, 1910).

Draba monoensis Rollins & R.A.Price

Brassicaceae Draba monoensis Rollins & R.A.Price -- Aliso 12: 22, fig. 1988 (GCI)

Astragalus monoensis Barneby

Fabaceae (Leguminosae) Astragalus monoensis Barneby -- Leafl. W. Bot. 4:55, fig, 7-15 (1944).

Astragalus monoensis Barneby, spec. nov. egregia sed affinitatis adhuc hinc stiputis connatis Chaetodontibus Gray inter quos ob tegumen obcompressum calycem muttoties superans anomata, vel Ocreatis Gray (Batidophacae ser. Humistratis Rydb.) a quibus tegumine subbitocutari graviter discrepat, illinc A. inyoensi Shetd. (Tii sp. Rydb.) qui ovario longe stipitato stiputisque disjunctis a nostra differt accedens, nec ulla cum specie (nisi cum A. striatifloro Jones, habitu notutisque compluribus haud absimiti sed quoad stytum exsertum carinamque acuminatam omnino singutari) arcte comparanda. Herba perennis humilis pulchella cryptophyta indumento e pilis patutis adscendentibusque brevibus plus minusve crispatis constituto prter petalos undique molliter villoso- vel subsericeo-cinerascens : caulibus numerosis e radicis verticalis graciliusculi pturicipitis collo subterraneo emissis, 1-2 dm. tongis, prostratis flexilibus sed haud abrupte flexuosis exangutatis hinc inde divaricatim ramosis, inferne filiformibus nudis, superne magis robustis et confertim fotiatis, tota tongitudine pube laxiuscule adscendenti indutis stiputis di fformibus, imis scariosis erectis vix 1 mm. tongis obtusis subperf ecte adnatis connatisque (ut cautem ocrea cuputiformi utrinque emarginata laxe vaginent) in medias summasque herbaceas ovato-acuminatas 2-3 mm. longas reflexas per tertiam partem tantum connatas breviusque adnatas gradatim accrescentibus, omnibus extus pubescentibus intus gtabris, imis nonnumquam demum gtabratis : foliis praesertim patulis, petioto gracili 3-6 (vel 12) mm. Longo incluso 1-3 cm. longis : foliolis (4 vel) 5-7-j ugis, approximatis vel foliorum summorum juniorumque valde congestis, oblongis obovatisve obtusis vel leviter retusis, 2-6 mm. longis, saepissime conduplicatis, manifeste falcatis, utrinque canescenti-villosulis, sursum secus rachin pauto decrescentibus : peduncutis adscendentibus rectis, folium suff ulcrans saltem breviter superantibus vel saepius eo dimidio tongioribus, in racemum 6-10-florum primo congestum subcapitutiformem demum laxiorem et ineunte fructu circa 1 cm. tongum abeuntibus bracteis subutatis herbaceis anguste scarioso-marginatis 1.5-3 mm. tongis pedicellum gracitem superantibus : floribus patentibus : calycis membranacei extus sericeovillosi tubo campanulato vet late tubuloso-campanulato 3-4 mm. longo, ad basin obliquam minutissime bibracteolato (sed bracteolae saepe in vestigia exigua aegre cernenda reductae sunt vet omnino deficiunt), ovario turgescenti mox rupto, dentibus subulatis acutiusculis circa 2 mm. longis, ciliatis, sinu lato obtusiusculo inter se separatis : petalis ex albo lutescentibus, vet vexillo dilute purpureo-striatuto carinaque apice macutata : vexillo obtanceolato-obovato profunde emarginato, exptanato circa 12 mm. Longo, in unguiculum Tatum sensim angustato, medium versus ad angutum fere rectum retroarcuato, marginibus patentibus vet leviter reflexis : atis subrectis, 10-11 mm. tongis, lamina obtanceolata obtusa auriculo parvo reflexo incluso 6 mm. Tonga carinae (7 vet) 8-9 mm. Tonga petalis secus margines tota fere longitudine connatis, unguiculis rectissimis circa 4.5 mm. longis, laminis oblique semiorbicularibus, inferne per annulum dimidium in apicem obtusum aequaliter arcuatis, superne levissime concavis subrectisve : legumine sessiti (vet ob pericarpium carnosulum in sicco contractum quasisubstipitato) chartaceo valde turgido, ambitu de visu ventrali ovoideo vet ovoideo-acuminato, de visu laterali lanceolato-lunato, basi abrupte obtuso vet truncato, apice acuto, valde fatcato et praeter rostrum acutum vet acuminatum vacuum compressum obcompresso, 1.5-2 cm. Longo, suturis ventrati per totam longitudinem acuta dorsali prominula sed depressa et infra medium legumen introversa ut septum praesertim incompletum 1-2.5 mm. altum suturam oppositam vix attingentem efformet, sectione compresse obcordata, 6-9 mm. lato, 3-5 mm. alto, valvulis pulchre reticulatis stramineopurpurascentibus pube brevissima strigoso-villosula indutis seminibus vatde compressis, 2-2.5 mm. longis, circa 2 mm. latis, ad hilum emarginatis, atrocastaneis.

CALIFORNIA : near Crestview, Mono Co., 10 Aug. 1938, J. T. Howell No. 14500. Type in Herb. Calif. Acad. Sci., No. 313539. Hills north of Mammoth, sandy plain, alt. 7900 ft., Peirson No. 6093. Valley north of Mammoth and east of Inyo Crater Lake, Mono Co., alt. 7500 ft., Ripley & Barneby No. 5833. Fig. 7-15.

Astragalus monoensis was first collected nearly twenty years ago by Mr. F. W. Peirson, to whom the writer is indebted for a portion of his original material. Since then it has been collected twice, always in the region immediately north of Mammoth, the three stations lying but a few miles apart. Mr. J. T. Howell, who kindly communicated the fine specimens selected as the type, alone has obtained the mature fruit.

The species is a remarkable one in that it exhibits a combination of characters not previously met with in North America. If the pod were considered the sole criterion of affinity in the genus — a course which Rydberg followed to unfortunate lengths — A. monoensis would be included quite naturally in Tium sensu Rydb., the falcate, obcompressed and imperfectly bilocular legume being not unlike that of A. salmonis Jones (though with the ventral, not the dorsal, suture concave) or that of A. inyoensis (though sessile). By the same system it might equally well be classified with certain varieties of A. lentiginosus Dougl. (Cystium Rydb.), though differing in the campanulate calyx. It departs, however, from all these species and their close allies in the connate stipules, a character which has proved of fundamental import almost throughout the genus, both in Asia as well as in the New World. Yet an attempt to attach the species to some natural group characterized by connate stipules is scarcely more successful. In general aspect A. monoensis is likely to recall A. sonorae Gray (sect. Ocreati Gray), but in that species, as well as its relatives, the stipules are scarious throughout, and the pod has no vestige of dissepiment. Perhaps more closely allied are the Chaetodontes (A. Austinae Gray etc.), but these are of rather different aspect, the long-villous, compressed pod is but little exserted from the calyx, and the keel of different shape. The species most closely resembling A. monoensis in the form of the stipules, in pubescence and in general habit is the singular A. striatiflorus Jones (sect. Cystiella Barneby), confined to a small area about Zion Park in southern Utah; but this species, with its attenuate keel, exserted style and membranous legume, seems otherwise remote. Yet again, in the general structure of the pod as well as in the obscurely bibracteolate calyx, A. monoensis may be linked with the very different A. iodanthus var. diaphanoides Barneby. It is probable that the species will eventually constitute a section apart.

In a narrow valley surrounded by pinewoods on the east slope of the Sierra Nevada north of Mammoth, A. monoensis is found in flat open places where the soil is composed of fine white gravel mixed with sand. Here, in a xerophytic association of low herbs which includes Hulsea vestita Gray, Oenothera xylocarpa Cov., and Eriogonum esmeraldense Wats., the species occurs locally but in some quantity, forming mats of silvery herbage closely pressed to the ground, or more often climbing up through the twigs of a dwarf Artemisia, where the whole plant becomes more lax and green. The numerous stems, filiform below but branched and stouter above, arise from the culticipital, subterranean crown of a slender taproot, rather in the manner of A. lentiginosus var. ineptus (Gray) Jones which grows nearby. The short leaves with their congested leaflets vested in villous hairs, the subcapitate racemes of proportionately large ochroleucous flowers, and the peculiar stipules are characteristic (Barneby, 1944).

Fabaceae (Leguminosae) Astragalus monoensis Barneby var. ravenii (Barneby) Isely -- Syst. Bot. 8(4): 423. 1983 (IK)

Lupinaster

Lupinus duranii Eastw.

Lupinus Durani Eastwood, spec. nov. Caudices multi, crassi, erecti, ex radice lignea, densissime tecti petiolis et stipulis prioribus et praesentibus, circa 1 dm. alti, dense tomentosi et vestiti pills albis adpressis et patentibus ; petiolis multo longioribus foliolis, 3-4 cm. longis, latioribus basi, stipulis 6 mm. longis, adnatis 2 mm., partibus disjunctis linearibus, acutis, 3 mm. longis, foliolis circa 6 vel 7, conduplicatis, 5-10 mm. longis, 4-5 mm. latis, anguste obovatis, infra vestitis similiter caudicibus, supra subglabris ; racemis folia superantibus, pedunculis aequilongis foliis, bracteis deciduis, pedicellis brevioribus calycibus, floribus circa 10 mm. longis, violaceis ; calyce basi saccato, labiis aequilongis, labio superiore bisecto segmentis ovatis, acutis, 3 mm. longis, sinu acuto, labio inferiore oblong.) veal ovato, bidentato ; vexillo aequilongo alis, 5 mm. lato, violaceo et fulvo-maculato, glabro, alis circa 3 mm. latis, carina glabra, tecta alis ; leguminibus 10-20 mm. longis, 5 mm. latis, seminibus circa 3, albis.

Type : Herb. Calif. Acad. Sci. No. 239801, collected on "Sand Flat," Mono County (near Mono Mills), California, at an elevation of 7800 ft. in open flats of pumice stone, July 15, 1932, by Victor Duran, No. 3343, in whose honor it is named. His specimens are always very good and from localities where but little collecting has been done. Mrs. Lester Rowntree collected the same species July 16, 1935, in earthquake fault region north of Mammoth. On road to June Lake, Mono County, California, Herb. Calif. Acad. Sci. No. 232190. A specimen collected by Mrs. Ynez Whilton Winblad July 28, 1938, at Lundy Lake, Mono County, at an elevation of 8500 ft. is the same, Herb. Calif. Acad. Sci. No. 273019.

This may be the same as L. breweri var. grandiflorus C. P. Smith, but in habit it does not suggest that trailing species and the flowers are much larger (Eastwood, 1940).

Lupinus monensis Eastwood

Lupinus monensis Eastwood, spec. nov. Caespitosus, nanus, erectus, circa 6-10 cm. altus, omnino vestitus albis villis adpressis et patentibus, argenteo-sericeis ; caudice ramoso ex radice lignea, basi dense folioso ; petiolis multo longioribus foliolis, 2-3 cm. longis, stipulis 1 cm. longis, adnatis 4-5 mm., partibus liberis linearibus attenuatis, foliolis 6 ad 8, oblanceolatis, circa 1 cm. longis, 3 mm. latis, mucronatis, supra vestitis villis adpressis, infra vestitis villis adpressis et patentibus ; pedunculis scaposis, aequalibus vel superantibus folia ; floribus violaceis, 1 cm. longis, verticillatis in racemis circa 3-4 cm. longis, 2 cm. latis, pedicellis brevioribus calycibus, bracteis lanceolatis, circa 8 mm. latis ; calyce 8 mm. longo, labio superiore bisecto basi 4 mm. lato, segmentis 3 mm. longis, sinu truncato, 3 mm. lato ; vexillo 8 mm. longo, circa 5 mm. lato, obovato, dorso villoso, medio albo ; alis anguste oblongis, 3 mm. latis, 8 mm. longis, stipite 2 mm. longo ; carina erecta obtusa, basi semi-sagittata, ciliata, stipite 2 mm. longo ; legumine 10-12 mm. longo, circa 8 mm. lato, seminibus 2 vel 3, albis.

Type : Herb. Calif. Acad. Sci. No. 272810, collected near Crestview, Mono County, California, Aug. 10, 1938, by John. Thomas Howell, No. 14498. It seems to be nearest to L. campbellae, but the flowers are longer, keel densely ciliate, and the pubescence more shaggy. A specimen collected by Anita Noldeke at Red Rock Canyon is the same, Herb. No. 272483; also one collected by Laura Lorraine, Aug. 6, 1938, "on bare pumice and obsidian slope, southeast slope of Mono Craters," Herb. No. 272484 (Eastwood, 1940).

Trifolium monoense Greene

Fabaceae Trifolium monoense Greene -- Erythea 2: 181. 1894 (GCI), Shockley #460, a synonym of Trifolium andersonii subsp. Monoense (Greene) J.M.Gillett., current name: Trifolium andersonii A. Gray subsp. Beatleyae J. M. Gillett

Fabaceae (Leguminosae) Lupinaster monoensis (Greene) Latsch. -- Zametki Sist. Geogr. Rast. 32: 22(1976). (IK)

Fabaceae Trifolium andersonii A.Gray var. monoense (Greene) Isely -- Brittonia 32(1): 55. 1980

Fabaceae (Leguminosae) Trifolium andersonii A.Gray subsp. Monoense (Greene) J.M.Gillett -- Canad. J. Bot. 50(10): 1997 (1972). (IK)

Mentzelia monoensis Brokaw & L.Hufford

Loasaceae Mentzelia monoensis Brokaw & L.Hufford -- Madroño 58(1): 57 (-63; figs. 1-3). 2011 [31 Aug 2011]

Oenothera heterochroma S. Watson var. monoensis Munz

Onagraceae Oenothera heterochroma S.Watson subsp. Monoensis (Munz) P.H.Raven -- Univ. Calif. Publ. Bot. 34: 113. 1962 (GCI)

Onagraceae Camissonia heterochroma subsp. Monoensis (Munz) P.H.Raven -- Brittonia 16: 282. 1964 (GCI)

Onagraceae Camissonia heterochroma (S.Watson) P.H.Raven var. monoensis (Munz) Cronquist -- Great Basin Naturalist 52: 76. 1992 (GCI)

Arctostaphylos uva-ursi (L.) Spreng. ssp. monoensis Roof

Ericaceae Arctostaphylos uva-ursi (L.) Spreng. subsp. monoensis Roof -- Changing Seasons 1(3): 7. 1980 (GCI)

Polemoniaceae

Aliciella monoensis J.M.Porter & A.G.Day

Polemoniaceae Aliciella monoensis J.M.Porter & A.G.Day -- Phytotaxa 15: 16 (-18; fig. 1). 2011 [28 Jan 2011]

Phacelia monoensis Halse

Hydrophyllaceae Phacelia monoensis Halse -- Madroño 28: 124 (-125). 1981 (GCI)

Castilleja rubida Piper var. monoensis (Jeps.) Edwin

Scrophulariaceae Castilleja rubida Piper var. monoensis (Jeps.) Edwin -- Leafl. W. Bot. 9: 72. 1959 (GCI)

Penstemon monoensis A.Heller

Scrophulariaceae Penstemon monoensis A.Heller -- Muhlenbergia 2: 246. 1906 (GCI)

Eriophyllum monoense Rydb.

The collection was described as Eriophyllum monoense Rydb. In N. Amer. Fl. 34(2): 87. 1915 [28 Jul 1915] (IK), as follows:

8. Eriophyllum monoense Rydberg, sp. Nov.

A perennial, with a decumbent cespitose base; stems about 1 dm. high, densely white- tomentose, very leafy; leaves obovate, 1 cm. long or less, with 3 rounded lobes at the apex, rather thick, densely white- tomentose on both sides; heads solitary at the ends of the stems; peduncles 3-4 cm. long; involucre nearly hemispheric, about , mm. high, 8-10 mm. broad, tomentose; bracts 8-10, elliptic, rounded or obtuse at the apex; ray-flowers 8-10; Hgules yellow, 7-8 mm. long, about 5 mm. wide; disk-corollas 4 mm. long; tube glandidar-hirsute, somewhat shorter than the puberulent throat ; achenes glabrous, striate, 3 mm. long; squamellae 8, unequal, the longer about 1.5 mm. long.

Type collected south of Mono Lake. July 8. 1863, Brewer 1823 (U. S. Nat. Herb. 323165).

Distribution: Middle Sierra Nevada, California.

The taxon was apparently first placed in synonomy with Eriophyllum lanatum var. integrifolium (Hook.) Smiley in Smiley's 1921 Flora of the Sierra Nevada of California, Univ. Calif. Publ. Bot. 9 378 1921.

2a. Eriophyllum lanatum var. integrifolium (Hook.), comb. Nov. (Tnchophyllum integrifolium Hook., FL Bor. Am., voL 1, p. 316. 1833. Bahia integrifoUa DC, Prodr., voL 5, p. 656. 1836. E. caespitosum var. integrifolium Graj, Proc. Am. Acad., vol. 19, p. 26. 1883. E. integrifolium Greene, Fl. Fran., p. 444. 1897. E, lutescena Bjdb., N. Am. FL, vol. 34, p. 87. 1915. E. monoense Rydb., kc.)

Type locality . — ' * With the preceding" on the * * Kooskoodcy River. ' '

Range. — Idaho to California; perhaps coextensive with the species.

Zone. — Transition to above timber line in rocky places.

Specimens eocamined. — Mt. Rose, 9,700 feet. Heller, August 28, 1911 ; Cisco Butte, 6,400 feet, Hall 8753a ; above Heather Lake, Tahoe, Setchell and Dobie, July 6-21, 1901 ; foot of Dicks Peak, Tahoe, 8,600 feet. Smiley 425 ; Sonora Pass, dry summit of the pass. Brewer 1900 ; Silver Mountain, Alpine County, 10-11,000 feet, Bolander 2688 ; Mt. Dana, 11,100 feet, Smiley 714; Cloud's Rest, Yosemite, Chesnut and Drew, July 10, 1889.

The variety differs from the species in having the leaves nearly entire, a few usually completely so in some plants, though, as a rule, the apex at least is distinctly lobed. The white woolly indument characteristic of this species is often so reduced in amount as to make the stems and leaves appear green rather than the typical cottony color. The several different aspects have been assigned specific values in the North American Flora.

The taxon was retained but reduced in rank to a variety by Jepson:

Eriophyllum lanatum (Pursh) J.Forbes var. monoense (Rydb.) Jeps. -- Man. Fl. Pl. Calif. [Jepson] 1119. 1925 (GCI)

The taxon was described in Abrams, Leroy. 1960. Illustrated Flora of the Pacific States, Volume 4. Stanford University Press.

Eriophyllum lanatum var. integrifolium (Hook.) Smiley, Univ. Calif. Pub. Rot. 9; 378. 1921. (Trichophylum integrifolium Hook. Fl. Bor. Amer. 1: 316. 1833; T. multiforum Nutt. Journ. Acad. Phila. 7: 35. 1834; Bahia gracilis Hook & Arn. Bot. Beechey 353. 1840; Eriophyllum caespitosum var. integrifolium A. Gray, Proc. Amer. Acad. 19: 26. 1883. in part; E. caespitosum var. leucophyllum A. Gray. Loc. Cit.. Not Bahia leucophylla DC. 1836; E. watsonii A. Gray, loc. Cit.; E. lutescens Rydb. N. Amer. Fl. 34: 87. 1915; E. monoense Rydb. Loc. Cit.; E. trichocarpum Rydb. Op. cit. 89; E. nevadense Gandoger, Bull. Soc. Bot. Fr. 65: 40. 1918.) Stems 1-2 dm. high, many, erect or decumbent from a woody base or short caudex, the herbage persistently tomentose, canescent or floccose; lower leaves entire or 3-5-toothed or -lobed at the apex, the stem leaves incised or pinnatifid above into 3 divisions; heads solitary or few on peduncles 3-10 cm. long; involucres 6-8 mm. high; rays 6-10 mm. long; achenes mostly hairy, clavate, the pappus present but extremely variable in this complex taxon. In the Pacific Northwest east of the Cascade Mountains in Oregon and Washington to Montana and Wyoming; in California it occurs in the northeastern counties southward in the Sierra Nevada, along the crest and on the eastern face, as far south as Tulare and Inyo Counties and also on the adjacent higher ranges in Nevada. Type locality: sources of the Columbia River.

Senecio douglasii var. monoensis (Greene) Jeps.

Asteraceae Senecio douglasii var. monoensis (Greene) Jeps. -- Man. Fl. Pl. Calif. [Jepson] 1149. 1925 (GCI)

Asteraceae Senecio flaccidus Less. Var. monoensis (Greene) B.L.Turner & T.M.Barkley -- Phytologia 69(1): 54 (1990):. (IK)

Senecio monoensis Greene

Senecio Monoensis. Woody at base, with many striate-angled stems decumbent, 1½ feet high, rather sparsely leafy, of a rather light green, the plant glabrous in all its parts; leaves made up of a filiform-linear rachis and few as narrowly linear remote acute segments: heads large, in a loose subcorymbose panicle; involucres broadly subcylindric, ½ inch high, notably calyculate-bracted at base, the bracts narrow, linear, acuminate: rays rather many and conspicuous, clear yellow.

White Mountains of Mono Co., Calif., on slate hills near Southern Belle Mine, A. A. Heller, n. 8330.

It turns out that the Southern Belle Mine is actually in Inyo County, being about 150 meters south of the Inyo-Mono county line.

Source Data About Other Non-Types or Non-Namesakes from Mono Lake or Mono County

Eriogonum umbellatum Torr.

Eriogonum umbellatum Torr. var. nevadense Gand.

Lupinus lepidus Douglas ex Lindl.

Lupinus lepidus has been shown to consist of varieties best described as ecotypes. Low genetic differentiation among varieties paired with high within population variation indicates that gene transfer between populations can be high, and that reproductive barriers between varieties either do not exist or have formed so recently as to not be detectable.

The varieties of L. lepidus can be viewed as divergent ecotypes, in which morphological differences may correlate with habitat differences, coupled with few barriers to reproduction between taxa (Clausen et al. 1939).

Clausen et al. (1939) accepted that ecotypes should be named as separate taxonomic entities and similarly it is recommended here that varieties of Lupinus lepidus retain recognition at the varietal rank.

Phlox stansburyi

Hesperochiron californicus (Benth.) S. Watson. California Monkey-Fiddle.

Sequuntur stirpes Californicae annis 1846 et 1847 lectae, imprimis circa Monterey et Sacramento.

Bentham (1857) placed Hartweg's collection in the Scrophulariaceae (p. 327):

1875 (367). Ourisia (Dichroma) californica, sp. N., foliis ad apicem rhizomatis brevis rosulatis petiolatis ovatis obtusis subintegerriniis ciliatis, pedicellis axillaribus scapiformibus unifloris folio brevioribus, calycis segmentis ovatis oblongisque ciliatis corollae tubo amplo brevioribus. —Rhizoma perpendiculare, breve. Folia circiter pollicem longa, basi cuneata, crassiuscula, supra pilis paucis conspersa, subtus glabra, petiolo subalato 2-5 lin. Longo basi dilatato. Pedicelli glabri, folio paullo breviores, ebracteati. Calyx 3 lin. Longus, segmento exteriore caeteris latiore, omnibus obtusis herbaceis praeter cilias marginales glabris. Corolla alba, 9 lin. Longa, late tubulosa, subinfundibuliformis, limbo obliquo subbilabiato, laciniis omnibus latis obtusis parum inaequalibus. Filamenta basi dilatata, ciliata. Stamina inclusa, didynama, adjecto quinto (in flore unico a me examinato an tamen constanter ?) caeteris paullo breviore sed pariter antherifero. —In uliginosis in montibus Sacramento.

Watson (1871) recognized that our plant was not a Scroph, but then placed it in Menyanthaceae the Buckbean family, as being similar to genus Villarsia.

Hesperochiron Californicus. (Ourisia Californica, Benth. Plant. Hartwg., p. 327.) More or less hirsute-pubescent; leaves clustered upon the summit of a short fusiform root, oblong or ovate, attenuate into a somewhat dilated fleshy petiole, obtuse, entire or obscurely repand-dentate, ciliate, 1-2' long; peduncles naked, 1-flowered, shorter than the leaves; calyx-segments 3" long, oblong, pubescent; corolla 5-8" long, more or less deeply cleft, the limb somewhat oblique, the tube and filaments somewhat hairy. —Pubescence rather variable, the surface of the leaves and sepals occasionally smoothish. An abnormal genus in some of its characters, its aestivation and the inequality of the corolla-lobes and stamens suggesting an affinity to the Scrophulariaceae, with which it was placed by Bentham without an examination of the ovary. It is, however, doubtless more nearly a Villarsia. Indeed, the V. pumila of Grisebach, (Hook. Flor. Bor.-Amer. 2. 70, t. 157; the corolla more open and somewhat rotate, the tube densely hairy within, and the leaves and calyx more glabrous,) is very similar and is probably either a second species or may prove to be but a form of the present one. From the Columbia River to the Sacramento, (1875 Hartweg, 379 Fremont, Cronkhite, and Lyall,) the " Snake Country," (Tolmie,) and near Carson City, (Anderson.) Found in the latter locality, in grassy meadows; 5,000 feet altitude; April. PLATE XXX. Fig. 1. A rather large plant; natural size. Figs. 2, 3. Flowers, different forms. Fig. 4. Flower, laid open; enlarged two diameters. Fig. 5. Stamen. Fig. 6. Ovary. Fig. 7. Mature capsule. Fig. 9. Seeds ; all enlarged four diameters. Fig. 8. Section of ovary ; enlarged eight diameters. (956.)

In the process, though, Watson published the genus name Hesperochiron in a footnote at the bottom of page 327:

HESPEROCHIRON. Calyx 5- (sometimes 6-7-) parted, the segments unequal. Corolla rather narrow campanulate, without either glands, corona or folds, the limb 5- (sometimes 6-7-) cleft, slightly bilablate and more or less spreading, the segments subequal, imbricate in aestivation. Stamens as many as the corolla-lobes, inserted at the base of the tube, unequal; filaments fleshy, attenuated upward; anthers oval, erect, cordate at base, 2-celled, cells laterally dehiscent. Ovary 1-celled, surrounded at base by 5 obscure glands, the numerous ovules attached in 2 rows to the sutural placentae. Styles united nearly to the apex, persistent; stigmas entire. Capsule 1-celled, loculicidally dehiscent, 2-valved, valves entire. Seeds wingless, ovate, roughened. —A perennial or biennial acaulescent herb,

I don't know yet who first placed the genus in Hydrophyllaceae.

6. Hesperochiron, S. Wats. Bot. 40 Parall. 281, t. 30. — Calycis lobi 5, rarius 6-7, ovati linearesve, inaequales. Corolla tubuloso-camapanulata v. subrotata, intus esquamata; lobi 5, rarius 6-7, lati, patentes, imbricati. Stamina 5, rarius 6-7, prope basin corollae affixa, filamentis filiformibus v. basin versus complanatis; antherae ovate. Ovarium 1-loculare, placentis 2 angustis parieti of affixis; stylus filiformis, apice breviter 2-fidis, stigmatibus parvis; ovula in quaque placenta ∞. Capsula 1-locularis, 2-valvis, valvis medio placentiferis. Semina ∞, angulato-subglobosa, rugosa. — Herbae nanae, acaules. Folia radicalia, petiolata, ovata v. oblonga, integerrima. Scapi v. pediunculi 1-flori, folia vix aequantes. Flores majusculi, albidi.

6. Hesperochiron, S. Wats. Bot. 40 Parall. 281, t. 30. - Calyx lobes 5, rarely 6-7, ovate linear, unequal. Corolla tubular-campanulate to subrotate, scale-less within; lobes 5, rarely 6-7, broad, patent, imbricate. Stamens 5, rarely 6-7, attached near the base of the corolla, the filaments filiform to base flattened; anthers ovate. Ovary 1-celled, 2-valves, parietal attachment, narrow; Stylus filiform, apex short, 2 stigmas, small; ova in each placenta many. Capsule 1-locule, 2-valves, septicidal. Seeds, many, angled-subglobose, wrinkled. - Herbs dwarf, acaulescent. Basal leaves, petiolate, ovate to oblong, entire. Scapes to peduncle 1-flowered, equalling the leaves. The flowers are larger, whitish.

Species 2, Americae borealis occidentalis incolae. A. Gray in Proc. Amer. Acad. x. 330. Huc pertinent Ourisia californica, Benth. Fl. Hartw. 327, et Villarsia pumila, Griseb. in Hook. Fl. Bor. Amer. ii. 70, t. i57 (Nicotiana nana, Lindl. Bot. Reg. t. 833). Genus anomalum tam a Scrophularineis Solanaceisve quam a Gentieneis pluribus notis distinctum, in omnibus cum Hydrophyllaceis melius convenire videtur.

Species: 2, inhabitants of western North America. A. Gray in Proc. Amer. Acad. x. 330. To this [genus] belongs Ourisia californica, Benth. Fl. Hartw. 327, and Villarsia pumila, Griseb. in Hook. Fl. Bor. Amer. ii. 70 t. I57 (Nicotiana nana, Lindl . Bot. Reg. t. 833). An anomalous genus in Scrophulariaceae and Solanaceae as well as Gentianaceae with several distinct characters that all seem to agree better with Hydrophyllaceae.

8. CAPNÒREA Raf. Dwarf stemless perennials. Leaves basal, oblong or spatulate, entire. Flowers singly on elongated pedicels from the axils of the leaves. Calyx 5-7-parted; lobes linear-lanceolate, often unequal. Corolla from campanulate or funnelform to saucer-shaped, deciduous, without appendages within. Stamens inserted in the base of the corolla, more or less hairy at the base. Ovary 1-celled, with 2 more or less intruding placentae; ovules 20 or more on each placenta; style 2-cleft at the apex. Capsule loculicidal, many-seeded. Seeds minutely reticulate. [Hesperochiron S. Wats.] Corolla saucer-shaped or rotate. Calyx almost glabrous, except the ciliate margins; leaves not distinctly venulose. ... 1. C. pumila. Calyx long-villous at the base; leaves distinctly venulose. ... 2. C. nervosa. Corolla campanulate or funnelform. Calyx-lobes oblong or lanceolate, obtuse. Corolla 12-15 mm. long; pubescence of the plant short. ... 3. C. Watsoniana. Corolla 7-10 mm long; pubescence longer on the peduncle; calyx densely villous. ... 4. C. incana. Calyx-lobes linear or linear-lanceolate, acute. ... 5. C. nana. 1. C. pumila (Dougl.) Greene. Leaves oblanceolate or elliptic, tapering into short petioles, 4-7 cm. long, glabrous except on the margins; scape 3-7 cm. long; calyx-lobes equal, lanceolate, 5-7 mm. long; corolla 10-15 mm. high. Hillsides: Wash. Ida.—Utah—n Calif. Submont. Je-Jl. 2. C. nervosa Greene. Leaves thin, petioled; blades elliptic, glabrous on both sides, sparingly silky-ciliate on the margin; calyx-tube densely villous; lobes lanceolate, long-ciliate on the margins; corolla fully 1 cm. high. Wet meadows: Ida. Submont. My. 3. C. Watsoniana Greene. Leaf-blades oblong or oblanceolate, obtuse, 3-7 cm. long, grayish pubescent, with short hairs; scape 2-5 cm. long; sepals oblong or elliptic, short-hairy; corolla lilac-purple, 12-15 mm. long. Gravelly soil: e Calif.—w Wyo. Submont. Ap-Je. 4. C. incana Greene. Leaf-blades elliptic or spatulate, 1-3 cm. long, canescent on both sides; scape 1-3 cm. long; calyx-lobes oblong or lanceolate, obtuse; corolla 7-10 mm. long; lobes rounded. Wet benches and flats: Mont.—Ida.—Wash. Submont. My-Je. 5. C. nana (Lindl.) Raf. Leaf-blades ovate-oblong or oval, strigose-pubes-cent, 2-5 cm. long; scape 4-7 cm. long; calyx-lobes 5-8 mm. long; corolla 10-17 mm. long; lobes ovate, acute. C. lasiantha Greene. C. macilenta Greene. Wet places and creek banks: Wash.—Ida.—Ore. Submont. Ap-Au.

Phacelia humilis Torr. & A. Gray

PHACELIA HUMILIS (n. sp.): annual, low, much branched from the base; leaves oblong, spatulate or lanceolate, all simple and entire, indistinctly veined, minutely hirsute-pubescent like the branches, and glandular dotted; racemes densely-flowered; segments of the calyx linear, obtuse, hispid, a little shorter than the (deep violet-colored) corolla; stamens exerted. Near the summit of the Sierra Nevada, California; June. A well-marked species, three or four inches high, somewhat cinereous, with a fine pubescence, except the inflorescence, and especially the calyx, which is hispid with rigid white hairs. Leaves an inch or less in length, shortpetioled. Corolla short, when expanded three lines in diameter; the base biplicate between the stamens. Filaments sparingly hispid above. Style glabrous. Ovules two in each cell. Capsule 2-3-seeded. This can hardly be the P. canescens of Nuttall, in P1. Gambell., which accords better with some states of P. circinata.

(Howell, 1943b) … in the key, uses "filaments long hairy" …

Munz (1965) does not use filament hairiness in his key, but does describe "… the fils. Pubescent …" in the description.

Taylor (2010) does not use filament hairiness in his key, nor does the character appear in his description.

TJM2 (Baldwin, 2012) in the key to Phacelia, couplet 18-18' says, "Filaments short-hairy," and describes P. humilis as "stamens short-hairy." No statement is made regarding hairiness for the varieties.

Phacelia humilis Torr. & A. Gray var. dudleyi J. T. Howell

TJM2 (Baldwin, 2012) describes "stamens 6-8 mm., clearly exserted"

Phacelia humilis Torr. & A. Gray var. humilis

TJM2 (Baldwin, 2012) describes "stamens 4-6 mm, barely exserted"

Mimulus glabratus var. utahensis

Section II. SIMIOLUS Greene

Mimulus, Sect.Simiolus Greene, in Bull. Calif. Acad. Sci. 1: 109. 1885. In this section, Dr. E. L. Greene placed all yellow-flowered species of Mimulus, and his diagnosis is sufficiently inclusive to apply to all. I follow Mrs. Grant in her restriction of the name to the present distinctive group of species although it must be noted that Greene's first species M. dentatus does not belong here; however, that species with its wide-throated corolla hardly agrees with the immediate group (defined as “corollas strongly bilabiate”) in which Greene placed it, so that, except on rigid grounds of text-priority, we may pass by it to the next-ensuing M. tilingii Regel and M. guttatus DC. It is the last wide- spread and abundant species for which the name Simiolus seems especially apt and which I consider as the logical type for the section.

Simiolus is a natural group of western North America and of Chile; it is highly variable and Dr. Greene and other workers have recognized many component species, while Mrs. Grant reduces them to sixteen.

Key to Species

A. Corolla 20-40 min. long, the throat much wider than high, the orifice nearly or quite closed by the uparching anterior lip; calyx-lobes all developed, at least the lateral acute; leaf blades ovate or oval, irregularly dentate; stems erect, or some- times slightly decumbent and rooting at the lower nodes . . . . . . . . . . . 2. M. guttatus

AA. Corolla 9-18 mm. long, the throat little wider than high, the orifice only partially closed; calyx-lobes usually blunter and the anterior tending to become obsolete; stems decumbent, extensively spreading and rooting at the nodes . . . . . . . . . . . . 3. M. glabratus

2. Mimulus guttatus Fischer

3. Mimulus glabratus Humboldt, Bonpland & Kunth (Map 23)

Like Mimulus guttatus and perhaps all species of Simiolus, the present species varies greatly in size, especially as to the leaves but also as to the calyx and corolla. The petioles may be well developed, or the leaf-blades may be nearly sessile. The leaf-blades may be acute or broadly rounded, dentate to entire, while the plants may be wholly glabrous or (as indicated by Dr. Grant) they may bear varying though slight amounts of pubescence. The species has a long meridian distribution from Manitoba to Chile, the typical subspecies growing at higher elevations from Mexico to Bolivia, replaced southward by the more pubescent subsp. parviflorus (Lindl.) Grant of middle Chile and Argentina, and northward by the following three subspecies.

The recognition of these Nearctic subspecies makes possible a reasonable geographic assignment of material, nearly all specimens falling satisfactorily into them. But intergradation is evident, and some specimens show characters of several subspecies; thus, E. J. Palmer 34325 and E. L. Reed 1819 from Alpine, Brewster County, Texas, classed below as subs. typicus, have uniform slightly denticulate leaf-blades as in subsp. fremontii; also, Pilsbry's plant from Buckeye, Arizona, classed below as subsp. fremontii, shows especially strongly dentate leaf-blades although the calyces are only 8-9 mm. long and the corolla only 9 mm. long. An actual study of all the biotypes of M. glabratus would evidently become an involved genetical problem. Of all its component elements the most remarkable appears to be the plant now being described as subsp. michiganensis; its large corollas and sinuate-dentate leaf-blades are unexpected in a form of very local occurrence within an area presumably heavily glaciated. Instead of intergradation to the neighboring subsp. fremontii, the plant has in accentuated degree characters elsewhere present only in subsp. typicus.

The Nearctic subspecies of Mimulus glabratus may be distinguished as follows:

A. Fruiting pedicels 10-20(-30) mm. long, twice as long as the subtending bracts; calyx 5-13 mm. long; leaf-blades usually at least denticulate.

B. Leaf-blades ovate or oval, denticulate or dentate; corolla usually over 12 mm. Long; calyx usually 10-13 mm. long; stems strongly ascending distally.

C. Corolla usually 12-16 mm. long, usually spotted within anteriorly; leaf- blades undulate-denticulate or -dentate . . . . . . . . . . . . 3a. M. g. typicus

CC. Corolla 15-22 mm. long, unspotted; leaf-blades sinuate-dentate. . . . . . . . . . . . . 3b. M. g. michiganensis

BB. Leaf-blades widely oval to suborbicular or uniform, undulate-denticulate to entire; corolla usually 8-12 mm. long, not spotted within anteriorly; calyx 5-10 mm. long; plant diffusely spreading . . . . . . . . . . . . 3c. M. g. fremontii

AA. Fruiting pedicels 25-50 mm. long, more than twice as long as. The subtending bracts; calyx 10-15 mm. long; leaf-blades oval to reniform, denticulate to entire. . . . . . . . . . . . . 3d. M. g. utahensis

3d. Mimulus glabrous utahensis Pennell, subsp. Nov.

Stem 4-5 dm. long, stout, finely pubescent at the nodes or more so on the distal portions of branches and on petioles. Leaf-blades cordate-orbicular, varying to oval or reniform, undulate-denticulate to usually nearly or quite entire. Pedicels 25-50 mm. long. Calyx becoming 12-15 min. long. Corolla usually 15-18 mm. long.

(Caulis robustior; folia cordato-orbicularia, saepe integra; pedicelli 25- 50 mm. Longi; calyx 12-15 mm. longus; corolla saepissime 15-18 mm. longa.)

Type, along brook, Preuss Lake, near Clay's Ranch, Millard County, Utah, collected in fruit and late flower August 29, 1919, by Ivar Tidestrom 11180; in Herb. Academy of Natural Sciences of Philadelphia.

Springheads and cool streams, western Colorado to western Nevada. In the Great Basin and the valley of the Colorado River.

Colorado. Montrose: Montrose, Shear 4807 (Y). Paradox, Walker 118 ®.

Utah. Preuss L., Tidestrom 11180 (Ph). Piute. Maryvale, Rydberg & Carlton 6989 (H, Y). Salt Lake: Salt Lake City, Pennell 5968 (Ph, Y). Utah: near Utah L., Bryan Exped. (M).

Nevada. Washoe: Reno, Brandegee (M, Ph).

• Rush Creek, Herbert L. Mason 13646. 9/4/1948 ( UC1191890)
• Mono Vista Spring, marshy area, Mono Vista Springs, northwest shore of Mono Lake Malcolm A. Nobs S. Galen Smith 1712. 9/1/1949 ( UC1191889)
• by lake G. Ledyard Stebbins 714. 4/19/1968 ( UC1462136)
• Old Marina, shallow water of springs dominated by Nasturtium officionale Mono Lake, just north of the town of Lee Vining and about 300 m east of Highway 395 at the site of the old marina Dean Wm. Taylor 11199. 6/27/1990 ( UC1755234)

Penstemon cinicola Keck

Penstemon cinicola Keck is described as follows (Clausen, Keck, and Heisey, 1940h, p. 294).

Penstemon cinicola Keck sp. nov.

Herba perenis glaber (saltem caules apud basem minute puberuli); caulibus herbaceis e basi suffrutescente erectus 15-35 cm. altis gracilibus foliosus; folis radicalibus fere nullis, caulinis linearibus moderate recurvis integerrimis 25-55 mm. longis 2-4 mm. latis, floralibus lanceolati-subulatis attenuatis; thyro stricto contracto; cymis subsessilibus multoflorus 3-6 geninis; calyce 1.6-2 mm, longo, lobis obovato-oblongis truncatis vel mucronatis late scarioso-marginatis; corolla purpureo-coerulea 7.5-9 mm. longa declinata, palato barbato; antheris glabris inclusis; filamento sterili apudapicem barbato.

Type, from among tussocks of grass and sedge on base volcanic sand, in openings of Pinus contorta forest just north of Lapine, Deschutes County Oregon, at 1285 m. (4225 ft.) elevation, June 23, 1935, Keck & Clausen 3690 (Dudley Herbarium of Stanford University); isotypes Academy of Natural Sciences, Philadelphia, and Carnegie Institution. This collection is tetraploid.

We refer the following collection from Klamath County, Oregon here: 20 miles south of Lapine on road to Silver Lake, Keck & Clausen 3692 (U. of Calif., Carnegie Inst., Phila., Pomona, Stanford, U. S. Nat. Herb.); Odell Lake, Howell 6911 (Calif. Acad., Carnegie Inst.); Annie Creek valley, near south entrance to Crater Lake National Park, Applegate 11113 (Carnegie Inst., Stanford).

Discussion of this new species in Clausen, Keck, and Heisey (1940h, p. 265).

Penstemon cinicola Keck is a newly described species (described [above]). It occurs only on volcanis ash in that great region of lava flows east of the Cascade Mountains in central Oregon and probably south to the lava beds of northern California. Its precise boundaries must be worked out with the aid of cytology, for certain outstanding characters, such as the diminuitive calyx, which were thought to mark this unit clearly have been discovered in the diploid forms of P. procerus from the borders of this region. Also, the peculiar habit of the plant, in failing to develop a basal rosette as do the meadow-dwelling members of P. procerus, must be verified a number of times by chromosome sounts before it can ne accepted as correlated with the tetraploid chromosome number. We have counted chromosomes in plants (thereby verified as procerus) from the meadows in the same local region in which P. cinicola is found growing on ash. Hybrids between the two have not been discovered.

Artemisia ludoviciana Nutt.

ARTEMISIA. L. (Wormwood, Southern-wood, &c.) Calix imbricated, scales rounded, connivent. Rays of the corolla none. Receptacle subvilluus, or nearly naked. Pappus none. Shrubby or herbaceous; leaves mostly multifid, flowers often racemose.

…

5. * ludoviciana. Stem simple and herbaceous; lower leaves incise, subpinnatifid, the upper lanceolate and entire, on both sides pubescent, beneath tomentose; flowers ovate, erect and sessile; calix pubescent, panicle simple — Hab. On the banks of the Missisippi, near St. Louis; also on the alluvial plains of the Missouri. Obs. Perennial. Stem about 2 feet high. Lower stem leaves lanceolate, irregularly and divaricately laciniate, segments entire, oblong-lanceolate and acute.

Artemisia ludoviciana Nutt. ssp. incompta (Nutt.) Keck

Artemisia incompta; herbaceous ; smooth, except the under surface of the leaves, which is a little tomentose; leaves almost simply pinnatifid, trifid or laciniate, sessile, the segments rather broad, linear and acute; flowers panicu- lated; capituli subglobose, pedicellate, erect; sepals ovate and scariose; florets numerous, smooth.

Hab. In the central chain of the Rocky Mountains, in Thornberg's Pass, near the great passage to the plains of the Oregon. At first sight it somewhat resembles some variety of A. vulgaris, but is very distinct. Remarkable for its smoothness. Height one to two feet; segments of the leaves a line wide.

(Clausen, Keck, and Heisey, 1940):

Representatives of Artemisia ludoviciana Nutt. are shown in figure 129. Plant 1324-3 on the left, from near Timberline station, is referable to A. ludoviciana ssp. incompta (Nutt.) Keck comb. nov. (A. incompta Nutt. Trans. Amer. Phil. Soc. II, 7:400, 1841). This subspecies has passed frequently under the name A. vulgaris ssp. discolor (Dougl.) H. & C., which is now thought referable to a more northern species, A. Michauxiana Bess. Subspecies incompta is the subalpine and alpine form of ludoviciana, distinguished by the deeply and twice cut divisions of the lower leaves. Much variation and doubtless more than a single ecotype are included in this subspecies, which occurs in the mountains of Montana and Idaho, southward to Utah, Nevada, and California.

Clausen, Keck, and Heisey (1940) continue with this discussion of ssp. ludoviciana

The remaining plants in figure 129 belong to A. ludoviciana ssp. typica Keck nom.nov. (A. ludoviciana Gen, 2: 143, 1918). We believe this form, except in the high mountains, where it is replaced by the preceding subspecies, and in the Northwest, where a larger-headed form intervenes, is the principal one found from the Great Basin to the Mississippi River. Rather locally it extends westward to the eastern flanks of the Cascades and Sierras. It is chiefly confined to the region bounded by the thirty-fifth and fifty-first parallels, but its exact boundaries fluctuate considerably. The Great Basin form of the subspecies came from Leevining, Mono County, California; the Rocky Mountain plants are from the east flank of Pikes Peak, Colorado. In 1328-1 the leaves are tomentose on both sides, and this form is widely known as A. gnaphalodes Nutt.; in 1329-1 the leaves are prominently discolored, almost glabrous above, and densely tomentose beneath, and this matches the type of A. ludoviciana Nutt. We consider these forms inseparable into natural subunits, for both sorts occur together intermittently almost throughout the range of the species and obviously intermix. In this instance these characters do not even mark ecotypical differences.

Madia glomerata Hook.

Madia gracilis (Sm.) Keck